Branched-chain ketoacids secreted by glioblastoma cells via MCT1 modulate macrophage phenotype

EMBO Rep. 2017 Dec;18(12):2172-2185. doi: 10.15252/embr.201744154. Epub 2017 Oct 24.

Abstract

Elevated amino acid catabolism is common to many cancers. Here, we show that glioblastoma are excreting large amounts of branched-chain ketoacids (BCKAs), metabolites of branched-chain amino acid (BCAA) catabolism. We show that efflux of BCKAs, as well as pyruvate, is mediated by the monocarboxylate transporter 1 (MCT1) in glioblastoma. MCT1 locates in close proximity to BCKA-generating branched-chain amino acid transaminase 1, suggesting possible functional interaction of the proteins. Using in vitro models, we demonstrate that tumor-excreted BCKAs can be taken up and re-aminated to BCAAs by tumor-associated macrophages. Furthermore, exposure to BCKAs reduced the phagocytic activity of macrophages. This study provides further evidence for the eminent role of BCAA catabolism in glioblastoma by demonstrating that tumor-excreted BCKAs might have a direct role in tumor immune suppression. Our data further suggest that the anti-proliferative effects of MCT1 knockdown observed by others might be related to the blocked excretion of BCKAs.

Keywords: BCAT1; MCT1; branched‐chain ketoacid; glioblastoma; phagocytosis.

MeSH terms

  • Amino Acids, Branched-Chain / metabolism*
  • Biological Transport
  • Cell Count
  • Cell Line, Tumor
  • Glioblastoma / immunology
  • Glioblastoma / physiopathology*
  • Humans
  • In Vitro Techniques
  • Macrophages / immunology
  • Macrophages / pathology
  • Macrophages / physiology*
  • Monocarboxylic Acid Transporters / antagonists & inhibitors
  • Monocarboxylic Acid Transporters / deficiency
  • Monocarboxylic Acid Transporters / genetics
  • Monocarboxylic Acid Transporters / metabolism*
  • Muscle Proteins / deficiency
  • Muscle Proteins / genetics
  • Phagocytosis
  • Phenotype
  • Pyruvic Acid / metabolism
  • Symporters / antagonists & inhibitors
  • Symporters / genetics
  • Symporters / metabolism*
  • Transaminases

Substances

  • Amino Acids, Branched-Chain
  • Monocarboxylic Acid Transporters
  • Muscle Proteins
  • SLC16A4 protein, human
  • Symporters
  • monocarboxylate transport protein 1
  • Pyruvic Acid
  • Transaminases