Inhibition of Asparagine-linked Glycosylation Participates in Hypoxia-induced Down-regulation of Cell-surface MICA Expression

Anticancer Res. 2018 Mar;38(3):1353-1359. doi: 10.21873/anticanres.12358.

Abstract

Background/aim: Hypoxia down-regulates the expression of cell surface major histocompatibility class I-related chain molecule A (MICA) without increasing its shedding. Recently, the inhibition of N-linked glycosylation was also shown to reduce the cell-surface expression of MICA. We investigated the participation of asparagine (Asn)-linked glycosylation in hypoxia-induced down-regulation of cell-surface MICA using osteosarcoma cells.

Materials and methods: The cell-surface expression and Asn-N-glycosylation of MICA were estimated by flow cytometry, and western blot analyses, respectively.

Results: Hypoxia reduced the expression of N-linked glycosylated MICA, as well as the ratio of N-linked glycosylated to non-glycosylated MICA. 2-Deoxy-D-glucose, which inhibits N-linked glycosylation, reduced the cell-surface expression of MICA under normoxia, while D-Mannose increased N-glycosylated MICA, increasing cell-surface MICA under hypoxia. Cells transfected with wild-type MICA expression vector expressed cell surface MICA more than those transfected with mutant MICA expression vectors designed for abrogation of N-linked glycosylation.

Conclusion: The inhibition of Asn-N-linked glycosylation participates in hypoxia-induced down-regulation of cell-surface expression of MICA.

Keywords: Hypoxia; MICA; NKG2D ligand; glycosylation; osteosarcoma.

MeSH terms

  • Amino Acid Sequence
  • Asparagine / genetics*
  • Asparagine / metabolism
  • Cell Hypoxia
  • Cell Line, Tumor
  • Cell Membrane / genetics
  • Cell Membrane / metabolism
  • Deoxyglucose / pharmacology
  • Down-Regulation*
  • Gene Expression Regulation, Neoplastic*
  • Glycosylation / drug effects
  • Histocompatibility Antigens Class I / genetics*
  • Histocompatibility Antigens Class I / metabolism
  • Humans
  • Osteosarcoma / genetics
  • Osteosarcoma / metabolism
  • Osteosarcoma / pathology

Substances

  • Histocompatibility Antigens Class I
  • MHC class I-related chain A
  • Asparagine
  • Deoxyglucose