Unique Francisella Phosphatidylethanolamine Acts as a Potent Anti-Inflammatory Lipid

J Innate Immun. 2018;10(4):291-305. doi: 10.1159/000489504. Epub 2018 Jul 3.

Abstract

Virulent Francisella tularensis subsp. tularensis (Ftt) is a dynamic, intracellular, bacterial pathogen. Its ability to evade and rapidly suppress host inflammatory responses is considered a key element for its profound virulence. We previously established that Ftt lipids play a role in inhibiting inflammation, but we did not determine the lipid species mediating this process. Here, we show that a unique, abundant, phosphatidylethanolamine (PE), present in Francisella, contributes to driving the suppression of inflammatory responses in human and mouse cells. Acyl chain lengths of this PE, C24: 0 and C10: 0, were key to the suppressive capabilities of Francisella PE. Addition of synthetic PE 24: 0-10: 0 resulted in the accumulation of PE in host cells for up to 24 h of incubation, and recapitulated the inhibition of inflammatory responses observed with native Ftt PE. Importantly, this novel PE significantly inhibited inflammatory responses driven by a medically and globally important flavivirus, dengue fever virus. Thus, targeting these lipids and/or the pathways that they manipulate represents a new strategy to combat immunosuppression engendered by Ftt, but they also show promise as a novel therapeutic intervention for significant viral infections.

Keywords: Bacteria; Dendritic cell; Inflammation; Macrophage; Phosphatidylethanolamine.

Publication types

  • Research Support, N.I.H., Intramural

MeSH terms

  • Animals
  • Anti-Inflammatory Agents / metabolism*
  • Bacterial Proteins / genetics
  • Cells, Cultured
  • Dendritic Cells / immunology*
  • Dendritic Cells / microbiology
  • Female
  • Francisella tularensis / physiology*
  • Humans
  • Immune Evasion
  • Inflammation / immunology*
  • Inflammation / microbiology
  • Macrophages / immunology*
  • Macrophages / microbiology
  • Mice
  • Mice, Inbred C57BL
  • Mutation / genetics
  • Phosphatidylethanolamines / metabolism*
  • Transferases (Other Substituted Phosphate Groups) / genetics
  • Tularemia / immunology*
  • Tularemia / microbiology

Substances

  • Anti-Inflammatory Agents
  • Bacterial Proteins
  • Phosphatidylethanolamines
  • phosphatidylethanolamine
  • Transferases (Other Substituted Phosphate Groups)
  • CDP-diacylglycerol choline O-phosphatidyltransferase