Enhanced photorespiration in transgenic rice over-expressing maize C4 phosphoenolpyruvate carboxylase gene contributes to alleviating low nitrogen stress

Plant Physiol Biochem. 2018 Sep:130:577-588. doi: 10.1016/j.plaphy.2018.08.013. Epub 2018 Aug 10.

Abstract

The objective of this study was to reveal the physiological and molecular mechanisms of low-nitrogen (N) tolerance in transgenic plant lines containing C4 phosphoenolpyruvate carboxylase (C4-PEPC) gene. The transgenic rice lines only over-expressing the maize C4-PEPC) (PC) and their untransformed wild type, Kitaake (WT), were used in this study. At different N levels, the dry weight, total N content, carbon and N levels, photorespiration-related enzymatic activities, gene expression levels and photorespiration-related product accumulations were measured, as were the transgenic lines' agronomic traits. The PC line, having lower total N and higher soluble sugar contents, was more tolerant to low-N stress than WT, which was consistent with its higher PEPC and lower N-assimilation-related enzyme activity levels. The photosynthetic parameters, enzymatic activity levels, transcripts and products related to photorespiration in PC were also greater than in WT under low-N conditions. This study showed that increased carbon levels in transgenic rice lines overexpressing C4-PEPC could help regulate the photorespiratory pathway under low-N conditions, conferring low-N tolerance and a higher grain yield per plant.

Keywords: Low-nitrogen stress; Phosphoenolpyruvate carboxylase; Photorespiration; Transgenic C(4)-PEPC-Overexpressing rice (Oryza sativa L.).

MeSH terms

  • Gene Expression Regulation, Plant
  • Genes, Plant
  • Nitrogen / deficiency*
  • Oryza / enzymology
  • Oryza / genetics
  • Oryza / metabolism
  • Phosphoenolpyruvate Carboxylase / genetics
  • Phosphoenolpyruvate Carboxylase / metabolism*
  • Photosynthesis*
  • Plants, Genetically Modified
  • Stress, Physiological
  • Zea mays / enzymology
  • Zea mays / genetics
  • Zea mays / metabolism

Substances

  • Phosphoenolpyruvate Carboxylase
  • Nitrogen