Yap/Taz mediates mTORC2-stimulated fibroblast activation and kidney fibrosis

J Biol Chem. 2018 Oct 19;293(42):16364-16375. doi: 10.1074/jbc.RA118.004073. Epub 2018 Aug 28.

Abstract

Our previously published study demonstrated that mammalian target of rapamycin complex 2 (mTORC2) signaling mediates TGFβ1-induced fibroblast activation. However, the underlying mechanisms for mTORC2 in stimulating fibroblast activation remain poorly understood. Here, we found that TGFβ1 could stimulate mTORC2 and Yap/Taz activation in NRK-49F cells. Blocking either mTORC2 or Yap/Taz signaling diminished TGFβ1-induced fibroblast activation. In addition, blockade of mTORC2 could down-regulate the expression of Yap/Taz, connective tissue growth factor (CTGF), and ankyrin repeat domain 1 (ANKRD1). Overexpression of constitutively active Taz (Taz-S89A) could restore fibroblast activation suppressed by PP242, an mTOR kinase inhibitor in NRK-49F cells. In mouse kidneys with unilateral ureter obstructive (UUO) nephropathy, both mTORC2 and Yap/Taz were activated in the interstitial myofibroblasts. Ablation of Rictor in fibroblasts/pericytes or blockade of mTOR signaling with PP242 attenuated Yap/Taz activation and UUO nephropathy in mice. Together, this study uncovers that targeting mTORC2 retards fibroblast activation and kidney fibrosis through suppressing Yap/Taz activation.

Keywords: Rictor; Yes-associated protein (YAP); cell signaling; fibroblast; fibrosis; kidney.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Acyltransferases
  • Adaptor Proteins, Signal Transducing / metabolism*
  • Animals
  • Cell Cycle Proteins
  • Cell Line
  • Fibroblasts / metabolism*
  • Fibrosis
  • Kidney Diseases / pathology*
  • Mechanistic Target of Rapamycin Complex 2 / physiology*
  • Mice
  • Phosphoproteins / metabolism*
  • Transcription Factors / metabolism*
  • Transforming Growth Factor beta1 / pharmacology
  • YAP-Signaling Proteins

Substances

  • Adaptor Proteins, Signal Transducing
  • Cell Cycle Proteins
  • Phosphoproteins
  • Tgfb1 protein, mouse
  • Transcription Factors
  • Transforming Growth Factor beta1
  • YAP-Signaling Proteins
  • Yap1 protein, mouse
  • Acyltransferases
  • tafazzin protein, mouse
  • Mechanistic Target of Rapamycin Complex 2