Global proteomic profiling of the uniquely human CHRFAM7A gene in transgenic mouse brain

Gene. 2019 Sep 25:714:143996. doi: 10.1016/j.gene.2019.143996. Epub 2019 Jul 23.

Abstract

The uniquely human α7-nAChR gene (CHRFAM7A) is evolved from the fusion of two partially duplicated genes, FAM7 and α7-nAChR gene (CHRNA7), and is inserted on same chromosome 15, 5' end of the CHRNA7 gene. Transcription of CHRFAM7A gene produces a 1256-bp open reading frame encoding dup-α7-nAChR, where a 27-aminoacid residues from FAM7 replaced the 146-aminoacid residues of the N-terminal extracellular ligand binding domain of α7-nAChR. In vitro, dup-α7-nAChR has been shown to form hetero-pentamer with α7-nAChR and dominant-negatively regulates the channel functions of α7-nAChR. However, the contribution of CHRFAM7A gene to the biology of α7-nAChR in the brain in vivo remains largely a matter of conjecture. CHRFAM7A transgenic mouse was created and differentially expressed proteins were profiled from the whole brain using iTRAQ-2D-LC-MS/MS proteomic technology. Proteins with a fold change of ≥1.2 or ≤0.83 and p < 0.05 were considered to be significant. Bioinformatics analysis showed that over-expression of the CHRFAM7A gene significantly modulated the proteins commonly involved in the signaling pathways of α7-nAChR-mediated neuropsychiatric disorders including Parkinson's disease, Alzheimer's disease, Huntington's disease, and alcoholism, suggesting that the CHRFAM7A gene contributes to the pathogenesis of neuropsychiatric disorders mostly likely through fine-tuning the functions of α7-nAChR in the brain.

Keywords: CHRFAM7A; CHRNA7; Human-specific gene; Neuropsychiatric disorders.

MeSH terms

  • Animals
  • Brain / metabolism
  • Chromatography, Liquid / methods
  • Chromosomes, Human, Pair 15 / genetics
  • Gene Expression Profiling / methods
  • Genes, Duplicate / genetics
  • Humans
  • Male
  • Mice
  • Mice, Inbred C57BL
  • Mice, Transgenic / genetics*
  • Proteomics / methods
  • Signal Transduction / genetics
  • Tandem Mass Spectrometry / methods
  • alpha7 Nicotinic Acetylcholine Receptor / genetics*

Substances

  • Chrna7 protein, human
  • alpha7 Nicotinic Acetylcholine Receptor