Transcriptional profiling indicates cAMP-driven reversal of HIV latency in monocytes occurs via transcription factor SP-1

Virology. 2020 Mar:542:40-53. doi: 10.1016/j.virol.2020.01.006. Epub 2020 Jan 18.

Abstract

Latent HIV reservoir is a major barrier to absolute HIV cure. Studies on latency reversal agents (LRA) have by far focused mainly on CD4+ T-lymphocytes, while myeloid reservoirs remain under-represented despite their persistence and key contribution to HIV pathogenesis. cAMP has been shown to increase HIV-1 transcription in latently-infected monocytes/macrophages. In this communication, we explored the potential of commercially available pharmacological drugs and phosphodiesterase inhibitors to reactivate HIV in latently-infected monocytic cell-line, U1. We showed that increased levels of intracellular cAMP reverse HIV latency in vitro, which is specific to cells of the myeloid lineage. High throughput RNA-seq analysis revealed that cAMP modulates transcriptional profile of latently HIV-infected cells and provides favourable cellular environment for HIV to produce viral proteins. This reactivation of latent HIV was inhibited by Mithramycin A, a selective Sp1 inhibitor, indicating that the reversal of HIV latency in monocytes is driven by transcription factor Sp1.

Keywords: Drug repurposing; HIV latency; Myeloid reservoir; Sp1; Transcriptional profiling; cAMP.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • 1-Methyl-3-isobutylxanthine / pharmacology
  • Colforsin / pharmacology
  • Cyclic AMP / metabolism
  • Gene Expression Profiling
  • Gene Regulatory Networks
  • Genome, Viral
  • HIV-1 / drug effects
  • HIV-1 / genetics*
  • HIV-1 / physiology*
  • Host Microbial Interactions / drug effects
  • Host Microbial Interactions / genetics
  • Host Microbial Interactions / physiology
  • Humans
  • Jurkat Cells
  • Models, Biological
  • Monocytes / drug effects
  • Monocytes / metabolism*
  • Monocytes / virology*
  • Phosphodiesterase Inhibitors / pharmacology
  • Sp1 Transcription Factor / metabolism*
  • U937 Cells
  • Virus Activation / drug effects
  • Virus Activation / genetics
  • Virus Activation / physiology
  • Virus Latency / drug effects
  • Virus Latency / genetics
  • Virus Latency / physiology*

Substances

  • Phosphodiesterase Inhibitors
  • Sp1 Transcription Factor
  • SP1 protein, human
  • Colforsin
  • Cyclic AMP
  • 1-Methyl-3-isobutylxanthine