Inhibition of miR-93 promotes interferon effector signaling to suppress influenza A infection by upregulating JAK1

Int Immunopharmacol. 2020 Sep:86:106754. doi: 10.1016/j.intimp.2020.106754. Epub 2020 Jul 8.

Abstract

Type I interferons play a critical role in host defense against influenza virus infection. Interferon cascade induces the expression of interferon-stimulated genes then subsequently promotes antiviral immune responses. The microRNAs are important regulators of innate immunity, but microRNAs-mediated regulation of interferon cascade during influenza infection remains to be fully identified. Here we found influenza A virus (IAV) infection significantly inhibited miR-93 expression in alveolar epithelial type II cells through RIG-I/JNK pathway. IAV-induced downregulation of miR-93 was found to upregulate JAK1, the target of miR-93, and then feedback promote antiviral innate response by facilitating IFN effector signaling. Importantly, in vivo administration of miR-93 antagomiR markedly suppressed IAV infection, protecting mice form IAVs -associated death. Hence, the inducible downregulation of miR-93 feedback suppress IAV infection by strengthening IFN-JAK-STAT pathway via JAK1 upregulation, and in vivo inhibition of miR-93 bears considerable therapeutic potential for suppressing IAV infection.

Keywords: Influenza A; Interferon-stimulated genes; JAK1; Type I inteferon; microRNA-93.

MeSH terms

  • Animals
  • Antagomirs / administration & dosage
  • Cell Line
  • Gene Expression Regulation
  • Humans
  • Influenza A virus / physiology*
  • Influenza, Human / immunology*
  • Interferons / genetics
  • Interferons / metabolism
  • Janus Kinase 1 / genetics
  • Janus Kinase 1 / metabolism
  • MAP Kinase Kinase 4 / metabolism
  • Male
  • Mice
  • Mice, Inbred C57BL
  • MicroRNAs / genetics*
  • Orthomyxoviridae Infections / immunology*
  • Receptors, Cell Surface / metabolism
  • Signal Transduction

Substances

  • Antagomirs
  • MicroRNAs
  • Mirn93 microRNA, mouse
  • Receptors, Cell Surface
  • Robo3 protein, mouse
  • Interferons
  • Janus Kinase 1
  • MAP Kinase Kinase 4