Metabolic Coordination of Cell Fate by α-Ketoglutarate-Dependent Dioxygenases

Trends Cell Biol. 2021 Jan;31(1):24-36. doi: 10.1016/j.tcb.2020.09.010. Epub 2020 Oct 19.

Abstract

Cell fate determination requires faithful execution of gene expression programs, which are increasingly recognized to respond to metabolic inputs. In particular, the family of α-ketoglutarate (αKG)-dependent dioxygenases, which include several chromatin-modifying enzymes, are emerging as key mediators of metabolic control of cell fate. αKG-dependent dioxygenases consume the metabolite αKG (also known as 2-oxoglutarate) as an obligate cosubstrate and are inhibited by succinate, fumarate, and 2-hydroxyglutarate. Here, we review the role of these metabolites in the control of dioxygenase activity and cell fate programs. We discuss the biochemical and transcriptional mechanisms enabling these metabolites to control cell fate and review evidence that nutrient availability shapes tissue-specific fate programs via αKG-dependent dioxygenases.

Keywords: 2-hydroxyglutarate; alpha-ketoglutarate; cell fate; chromatin modifications; succinate; αKG-dependent dioxygenases.

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't
  • Review

MeSH terms

  • Alpha-Ketoglutarate-Dependent Dioxygenase FTO / metabolism*
  • Animals
  • Cell Lineage* / genetics
  • Enhancer Elements, Genetic / genetics
  • Humans
  • Mutation / genetics
  • Neoplasms / enzymology
  • Neoplasms / genetics
  • Transcription Factors / metabolism

Substances

  • Transcription Factors
  • Alpha-Ketoglutarate-Dependent Dioxygenase FTO