1 Arthropod-Borne Animal Diseases Research Unit, United States Department of Agriculture (USDA), Agricultural Research Service, Manhattan, KS, United States.
3 French Agency for Food, Environmental and Occupational Health & Safety (ANSES), Maisons Alfort, France.
4 Department of Diagnostic Medicine/Pathobiology and Center of Excellence for Emerging and Zoonotic Animal Diseases, College of Veterinary Medicine, Kansas State University, Manhattan, KS, United States.