Brucella-Induced Downregulation of lncRNA Gm28309 Triggers Macrophages Inflammatory Response Through the miR-3068-5p/NF-κB Pathway

Front Immunol. 2020 Dec 22:11:581517. doi: 10.3389/fimmu.2020.581517. eCollection 2020.

Abstract

Objectives: The underlying mechanism of the inflammatory response against Brucellosis caused by Brucella remains poorly understood. This study aimed to determine the role of long non-coding RNAs (lncRNAs) in regulating of inflammatory and anti-Brucella responses.

Materials and methods: Microarray analysis was performed to detect differentially expressed lncRNAs in THP-1 cells infected with an S2308 Brucella strain. The candidate lncRNAs were screened using bioinformatic analysis and siRNAs; bioinformatic prediction and luciferase reporter assay were also conducted, while inflammatory responses was assessed using RT-qPCR, western blot, immunofluorescence, ELISA, HE, and immunohistochemistry.

Results: The lncRNA Gm28309 was identified to be involved in regulating inflammation induced by Brucella. Gm28309, localized in the cytoplasm, was down-expressed in RAW264.7 cells infected with S2308. Overexpression of Gm28309 or inhibition of miR-3068-5p repressed p65 phosphorylation and reduced NLRP3 inflammasome and IL-1β and IL-18 secretion. Mechanistically, Gm28309 acted as a ceRNA of miR-3068-5p to activate NF-κB pathway by targeting κB-Ras2, an inhibitor of NF-κB signaling. Moreover, the number of intracellular Brucella was higher when Gm28309 was overexpressed or when miR-3068-5p or p65 was inhibited. However, these effects were reversed by the miR-3068-5p mimic.

Conclusions: Our study demonstrates, for the first time, that LncRNAs are involved in regulating immune responses during Brucella infection, and Gm28309, an lncRNA, plays a crucial role in activating NF-κB/NLRP3 inflammasome signaling pathway.

Keywords: Brucella; LncRNA Gm28309; NF-κB; inflammation; miR-3068-5p.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Brucella / immunology*
  • Cell Line
  • Down-Regulation / immunology*
  • Female
  • Humans
  • Inflammation / immunology*
  • Interleukin-18 / immunology
  • Interleukin-1beta / immunology
  • Macrophages / immunology*
  • Mice
  • Mice, Inbred BALB C
  • MicroRNAs / immunology*
  • NF-kappa B / immunology*
  • RAW 264.7 Cells
  • RNA, Long Noncoding / immunology*
  • Signal Transduction / immunology
  • THP-1 Cells

Substances

  • Interleukin-18
  • Interleukin-1beta
  • MicroRNAs
  • NF-kappa B
  • RNA, Long Noncoding