1 Evolution and Adaptation Group, German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig, Puschstraße 4, 04103 Leipzig, Germany.
2 Department of Theoretical and Computational Ecology, Institute for Biodiversity and Ecosystem Dynamics (IBED), University of Amsterdam, PO Box 94240, Amsterdam, The Netherlands.
3 Department of Biological Sciences, University of Bergen, PO Box 7803, 5020, Bergen, Norway.
4 School of Biological and Environmental Sciences, Liverpool John Moores University, Byrom Street, Liverpool L3 3AF, UK.