hnRNPA1 enhances FOXP3 stability to promote the differentiation and functions of regulatory T cells

FEBS Lett. 2021 Jul;595(14):1962-1974. doi: 10.1002/1873-3468.14142. Epub 2021 Jun 22.

Abstract

Regulatory T cells (Tregs) are indispensable for the maintenance of immunological self-tolerance and homeostasis. Heterogeneous nuclear ribonucleoprotein A1 (hnRNPA1) is required for optimal Treg induction. Here, we reveal that human-induced Tregs (iTregs) lacking hnRNPA1 show reduced expression of the transcription factor FOXP3, increased ubiquitination level of FOXP3, and impaired suppressive abilities. Human naïve CD4 T cells with hnRNPA1 knockdown show a decreased Treg differentiation ratio. hnRNPA1 could interact with FOXP3 as well as with the E3 ligase Stub1. The phosphorylation at hnRNPA1 S199 could increase both interactions. The overexpression of FOXP3 in Tregs containing shhnRNPA1 could not recover the phenotype caused by hnRNPA1 knockdown. Therefore, there might be multiple essential pathways regulated by hnRNPA1 in Tregs. In conclusion, we present a new role of hnRNPA1 in promoting Treg function, indicating it as a promising target for tumor therapies.

Keywords: FOXP3; Stub1; Tregs; hnRNPA1; ubiquitination.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Cell Differentiation
  • Forkhead Transcription Factors / genetics*
  • Forkhead Transcription Factors / immunology
  • Gene Expression Regulation
  • HEK293 Cells
  • Heterogeneous Nuclear Ribonucleoprotein A1 / antagonists & inhibitors
  • Heterogeneous Nuclear Ribonucleoprotein A1 / genetics*
  • Heterogeneous Nuclear Ribonucleoprotein A1 / immunology
  • Homeostasis / immunology
  • Humans
  • Phosphorylation
  • Primary Cell Culture
  • Protein Binding
  • Protein Stability
  • RNA, Small Interfering / genetics
  • RNA, Small Interfering / metabolism
  • Self Tolerance / genetics
  • Signal Transduction
  • T-Lymphocytes, Cytotoxic / cytology
  • T-Lymphocytes, Cytotoxic / immunology*
  • T-Lymphocytes, Regulatory / cytology
  • T-Lymphocytes, Regulatory / immunology*
  • Ubiquitin-Protein Ligases / genetics*
  • Ubiquitin-Protein Ligases / immunology
  • Ubiquitination

Substances

  • FOXP3 protein, human
  • Forkhead Transcription Factors
  • Heterogeneous Nuclear Ribonucleoprotein A1
  • RNA, Small Interfering
  • hnRNPA1 protein, human
  • STUB1 protein, human
  • Ubiquitin-Protein Ligases