UPRmt activation protects against MPP+-induced toxicity in a cell culture model of Parkinson's disease

Biochem Biophys Res Commun. 2021 Sep 10:569:17-22. doi: 10.1016/j.bbrc.2021.06.079. Epub 2021 Jun 30.

Abstract

The pathogenesis of Parkinson's disease (PD) remains elusive, but mitochondrial dysfunction is believed to be one crucial step in its pathogenesis. The mitochondrial unfolded protein response (UPRmt) is an important mitochondrial quality control strategy that maintains mitochondrial function in response to disturbances of mitochondrial protein homeostasis. Activation of the UPRmt and the beneficial effect of rescuing mitochondrial proteostasis have been reported in several genetic models of PD. However, the pathogenic relevance of the UPRmt in idiopathic PD is unknown. The present study examined the link between the UPRmt and mitochondrial dysfunction in 1-methyl-4-phenylpyridinium (MPP+)-treated SH-SY5Y cells. Treatment with MPP + induced activation of the UPRmt, reflected by an increase in the expression of UPRmt-related chaperones, proteases, and transcription mediators. UPRmt activation that was induced by overexpressing mutant ornithine transcarbamylase significantly reduced the production of mitochondrial reactive oxygen species (ROS) and improved cell survival in SH-SY5Y cells following MPP+ treatment. Moreover, the overexpression of activating transcription factor 5 (mammalian UPRmt transcription factor) conferred protection against MPP+-induced ROS production and against cell death in SH-SY5Y cells. Overall, our results demonstrate the beneficial effect of UPRmt activation in MPP + -treated cells, shedding new light on the mechanism of mitochondrial dysfunction in the pathogenesis of PD.

Keywords: ATF5; MPP(+); Parkinson's disease; UPR(mt).

Publication types

  • Research Support, N.I.H., Extramural
  • Research Support, Non-U.S. Gov't

MeSH terms

  • 1-Methyl-4-phenylpyridinium / pharmacology*
  • Activating Transcription Factors / genetics
  • Activating Transcription Factors / metabolism
  • Cell Line, Tumor
  • Cell Survival / drug effects
  • Cell Survival / genetics
  • Chaperonin 60 / genetics
  • Chaperonin 60 / metabolism
  • Endopeptidase Clp / genetics
  • Endopeptidase Clp / metabolism
  • Gene Expression / drug effects
  • Humans
  • Mitochondria / metabolism*
  • Mitochondrial Proteins / genetics
  • Mitochondrial Proteins / metabolism
  • Models, Biological*
  • Parkinson Disease / genetics
  • Parkinson Disease / metabolism*
  • Protective Agents / metabolism
  • Reactive Oxygen Species / metabolism
  • Ubiquitins / genetics
  • Ubiquitins / metabolism
  • Unfolded Protein Response / drug effects*
  • Unfolded Protein Response / genetics

Substances

  • ATF5 protein, human
  • Activating Transcription Factors
  • Chaperonin 60
  • HSPD1 protein, human
  • Mitochondrial Proteins
  • Protective Agents
  • Reactive Oxygen Species
  • UBL5 protein, human
  • Ubiquitins
  • ClpP protein, human
  • Endopeptidase Clp
  • 1-Methyl-4-phenylpyridinium