Abstract
The pandemic caused by severe acute respiratory syndrome coronavirus 2 (SARS-CoV-2) has resulted in over 100 million infections and millions of deaths. Effective vaccines remain the best hope of curtailing SARS-CoV-2 transmission, morbidity, and mortality. The vaccines in current use require cold storage and sophisticated manufacturing capacity, which complicates their distribution, especially in less developed countries. We report the development of a candidate SARS-CoV-2 vaccine that is purely protein based and directly targets antigen-presenting cells. It consists of the SARS-CoV-2 Spike receptor-binding domain (SpikeRBD) fused to an alpaca-derived nanobody that recognizes class II major histocompatibility complex antigens (VHHMHCII). This vaccine elicits robust humoral and cellular immunity against SARS-CoV-2 and its variants. Both young and aged mice immunized with two doses of VHHMHCII-SpikeRBD elicit high-titer binding and neutralizing antibodies. Immunization also induces strong cellular immunity, including a robust CD8 T cell response. VHHMHCII-SpikeRBD is stable for at least 7 d at room temperature and can be lyophilized without loss of efficacy.
Keywords:
COVID-19; nanobody; vaccine.
Copyright © 2021 the Author(s). Published by PNAS.
Publication types
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Research Support, N.I.H., Extramural
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Research Support, Non-U.S. Gov't
MeSH terms
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Amino Acid Sequence
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Animals
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Antibodies, Neutralizing / biosynthesis
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Antibodies, Viral / biosynthesis
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Antigen-Presenting Cells / immunology
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CD8-Positive T-Lymphocytes / immunology
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COVID-19 / epidemiology
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COVID-19 / immunology*
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COVID-19 / prevention & control*
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COVID-19 Vaccines / administration & dosage
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COVID-19 Vaccines / immunology*
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COVID-19 Vaccines / pharmacology*
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Camelids, New World / immunology
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Female
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Histocompatibility Antigens Class II / immunology
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Humans
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Immunity, Cellular
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Immunity, Humoral
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Immunization, Secondary
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Mice
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Mice, Inbred BALB C
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Mice, Inbred C57BL
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Mice, Transgenic
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Pandemics* / prevention & control
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Recombinant Fusion Proteins / administration & dosage
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Recombinant Fusion Proteins / genetics
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Recombinant Fusion Proteins / immunology
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SARS-CoV-2 / genetics
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SARS-CoV-2 / immunology*
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Single-Domain Antibodies / administration & dosage
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Single-Domain Antibodies / immunology
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Spike Glycoprotein, Coronavirus / administration & dosage
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Spike Glycoprotein, Coronavirus / genetics
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Spike Glycoprotein, Coronavirus / immunology
Substances
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Antibodies, Neutralizing
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Antibodies, Viral
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COVID-19 Vaccines
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Histocompatibility Antigens Class II
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Recombinant Fusion Proteins
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Single-Domain Antibodies
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Spike Glycoprotein, Coronavirus
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spike protein, SARS-CoV-2