Bile acids promote the caveolae-associated entry of swine acute diarrhea syndrome coronavirus in porcine intestinal enteroids

PLoS Pathog. 2022 Jun 13;18(6):e1010620. doi: 10.1371/journal.ppat.1010620. eCollection 2022 Jun.

Abstract

Intestinal microbial metabolites have been increasingly recognized as important regulators of enteric viral infection. However, very little information is available about which specific microbiota-derived metabolites are crucial for swine enteric coronavirus (SECoV) infection in vivo. Using swine acute diarrhea syndrome (SADS)-CoV as a model, we were able to identify a greatly altered bile acid (BA) profile in the small intestine of infected piglets by untargeted metabolomic analysis. Using a newly established ex vivo model-the stem cell-derived porcine intestinal enteroid (PIE) culture-we demonstrated that certain BAs, cholic acid (CA) in particular, enhance SADS-CoV replication by acting on PIEs at the early phase of infection. We ruled out the possibility that CA exerts an augmenting effect on viral replication through classic farnesoid X receptor or Takeda G protein-coupled receptor 5 signaling, innate immune suppression or viral attachment. BA induced multiple cellular responses including rapid changes in caveolae-mediated endocytosis, endosomal acidification and dynamics of the endosomal/lysosomal system that are critical for SADS-CoV replication. Thus, our findings shed light on how SECoVs exploit microbiome-derived metabolite BAs to swiftly establish viral infection and accelerate replication within the intestinal microenvironment.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Alphacoronavirus* / physiology
  • Animals
  • Bile Acids and Salts
  • Caveolae
  • Coronavirus Infections*
  • Diarrhea
  • Swine
  • Swine Diseases*

Substances

  • Bile Acids and Salts

Supplementary concepts

  • Swine acute diarrhea syndrome coronavirus

Grants and funding

This research was funded by the National Key Research and Development Program of China (2021YFD1801103) to YWH, and supported by the National Natural Science Foundation of China (No. U21A20261 and No. 32172864) to SJZ and by the Laboratory of Lingnan Modern Agriculture Project (NG2022001) to YWH. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.