Binding of heparan sulfate to human cystatin C modulates inhibition of cathepsin L: Putative consequences in mucopolysaccharidosis

Carbohydr Polym. 2022 Oct 1:293:119734. doi: 10.1016/j.carbpol.2022.119734. Epub 2022 Jun 18.

Abstract

Mucopolysaccharidoses (MPS) are a group of rare lysosomal storage diseases characterized by glycosaminoglycan (GAG) accumulation causing progressive multi-organs dysfunction and ultimately severe cardio-respiratory damages. Human cystatin C (hCC), a potent inhibitor of cysteine cathepsins, plays an important role in respiratory diseases. However, its regulation remained unknown in MPS. Herein, elevated hCC levels were measured in respiratory specimens from MPS-I, -II, and -III patients and were significantly correlated with severe respiratory symptoms (rs = 0.7173). Heparan sulfate (HS), a prominent GAG, dampened its inhibitory activity toward cathepsin L in a dose-dependent manner. HS and HS-oligosaccharides bound tightly hCC, in combination with a secondary structure rearrangement. Molecular modeling studies identified three HS binding regions in hCC, including the N-terminus, which is crucial in the inhibition of cathepsins. Impairment of inhibitory potential of hCC may reflect abnormal regulation of proteolytic activity of cathepsin L in lung, ultimately contributing to the severity of MPS.

Keywords: Glycosaminoglycan; Inhibitor; Lung; Molecular dynamic; Protease.

MeSH terms

  • Cathepsin L
  • Cystatin C*
  • Glycosaminoglycans / metabolism
  • Heparitin Sulfate
  • Humans
  • Mucopolysaccharidoses*

Substances

  • Cystatin C
  • Glycosaminoglycans
  • Heparitin Sulfate
  • Cathepsin L