1 Key Laboratory of Tropical Marine Bio-resources and Ecology, Guangdong Provincial Key Laboratory of Applied Marine Biology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China.
2 World Wide Fund for Nature Hong Kong, Hong Kong, China.
3 ARC Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, QLD, Australia.
4 Sanya National Marine Ecosystem Research Station, Tropical Marine Biological Research Station in Hainan, Chinese Academy of Sciences, Sanya, China.