1 CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla, Yunnan, 666303, China.
2 Yuanjiang Savanna Ecosystem Research Station, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Yuanjiang, Yunnan, 653300, China.
3 University of Chinese Academy of Sciences, Beijing, 100049, China.
4 School of Biology and Ecology, University of Maine, Orono, ME, 04469, USA.
5 Climate Change Institute, University of Maine, Orono, ME, 04469, USA.