The Sm core protein SmEb regulates salt stress responses through maintaining proper splicing of RCD1 pre-mRNA in Arabidopsis

J Integr Plant Biol. 2023 Jun;65(6):1383-1393. doi: 10.1111/jipb.13457. Epub 2023 Feb 27.

Abstract

Salt stress adversely impacts crop production. Several spliceosome components have been implicated in regulating salt stress responses in plants, however, the underlying molecular basis is still unclear. Here we report that the spliceosomal core protein SmEb is essential to salt tolerance in Arabidopsis. Transcriptome analysis showed that SmEb modulates alternative splicing of hundreds of pre-mRNAs in plant response to salt stress. Further study revealed that SmEb is crucial in maintaining proper ratio of two RCD1 splicing variants (RCD1.1/RCD1.2) important for salt stress response. In addition, RCD1.1 but not RCD1.2 is able to interact with the stress regulators and attenuates salt-sensitivity by decreasing salt-induced cell death in smeb-1 mutant. Together, our findings uncovered the essential role of SmEb in the regulation of alternative pre-mRNA splicing in salt stress response.

Keywords: RCD1; SmEb; alternative splicing; salt stress; spliceosome.

MeSH terms

  • Alternative Splicing / genetics
  • Arabidopsis Proteins* / metabolism
  • Arabidopsis* / metabolism
  • Gene Expression Regulation, Plant / genetics
  • RNA Precursors / genetics
  • RNA Precursors / metabolism
  • Salt Tolerance / genetics
  • Stress, Physiological / genetics

Substances

  • Arabidopsis Proteins
  • RNA Precursors
  • RCD1 protein, Arabidopsis