1 Department of Systematic & Evolutionary Botany, University of Zurich, Zollikerstrasse 107, 8008 Zurich, Switzerland University of Zurich Zurich Switzerland.
2 Universidade Estadual de Feira de Santana, Departamento de Cièncias Biológicas, Av. Transnordestina s.n., Novo Horizonte, Feira de Santana, BA, 44036-900, Brazil Universidade Estadual de Feira de Santana Feira de Santana Brazil.
3 Accelerated Taxonomy Department, Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3AE, UK Accelerated Taxonomy Department, Royal Botanic Gardens Richmond United Kingdom.