EBF1-JAK2 inhibits the PAX5 function through physical interaction with PAX5 and kinase activity

Int J Hematol. 2023 Jul;118(1):65-74. doi: 10.1007/s12185-023-03585-z. Epub 2023 May 7.

Abstract

Gene aberrations of B-cell regulators and growth signal components such as the JAK-STAT pathway are frequently found in B-cell acute lymphoblastic leukemia (B-ALL). EBF1 is a B-cell regulator that regulates the expression of PAX5 and co-operates with PAX5 to regulate B-cell differentiation. Here, we analyzed the function of the fusion protein of EBF1 and JAK2, EBF1-JAK2 (E-J). E-J caused constitutive activation of JAK-STAT and MAPK pathways and induced autonomous cell growth in a cytokine-dependent cell line. E-J did not affect the transcriptional activity of EBF1 but inhibited that of PAX5. Both the physical interaction of E-J with PAX5 and kinase activity of E-J were required for E-J to inhibit PAX5 function, although the detailed mechanism of inhibition remains unclear. Importantly, gene set enrichment analysis using the results of our previous RNA-seq data of 323 primary BCR-ABL1-negative ALL samples demonstrated repression of the transcriptional target genes of PAX5 in E-J-positive ALL cells, which suggests that E-J also inhibited PAX5 function in ALL cells. Our results shed new light on the mechanisms of differentiation block by kinase fusion proteins.

Keywords: Acute lymphoblastic leukemia; EBF1–JAK2; PAX5; Phosphorylation; RNA-seq.

MeSH terms

  • Cell Line
  • Humans
  • Janus Kinase 2 / genetics
  • Janus Kinase 2 / metabolism
  • Janus Kinases* / metabolism
  • PAX5 Transcription Factor / genetics
  • PAX5 Transcription Factor / metabolism
  • STAT Transcription Factors* / metabolism
  • Signal Transduction
  • Trans-Activators / genetics
  • Trans-Activators / metabolism
  • Transcription Factors / metabolism

Substances

  • Janus Kinases
  • STAT Transcription Factors
  • Transcription Factors
  • PAX5 Transcription Factor
  • EBF1 protein, human
  • Trans-Activators
  • PAX5 protein, human
  • JAK2 protein, human
  • Janus Kinase 2