1 State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources, Alpine Paleoecology and Human Adaptation Group, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China.
2 Xizang Institute for Conservation and Research of Cultural Relics, Lhasa 850000, China.
3 School of Geography and Remote Sensing, Guangzhou University, Guangzhou 510006, China.
4 Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China.
5 State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources, Alpine Paleoecology and Human Adaptation Group, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China. Electronic address: [email protected].