Global transcriptome analysis reveals Salmonella Typhimurium employs nitrate metabolism to combat bile stress

FEBS Lett. 2024 Jul;598(13):1605-1619. doi: 10.1002/1873-3468.14853. Epub 2024 Mar 19.

Abstract

Salmonella Typhimurium is an enteric pathogen that is highly tolerant to bile. Next-generation mRNA sequencing was performed to analyze the adaptive responses to bile in two S. Typhimurium strains: wild type (WT) and a mutant lacking cold shock protein E (ΔcspE). CspE is an RNA chaperone which is crucial for survival of S. Typhimurium during bile stress. This study identifies transcriptional responses in bile-tolerant WT and bile-sensitive ΔcspE. Upregulation of several genes involved in nitrate metabolism was observed, including fnr, a global regulator of nitrate metabolism. Notably, Δfnr was susceptible to bile stress. Also, complementation with fnr lowered reactive oxygen species and enhanced the survival of bile-sensitive ΔcspE. Importantly, intracellular nitrite amounts were highly induced in bile-treated WT compared to ΔcspE. Also, the WT strain pre-treated with nitrate displayed better growth with bile. These results demonstrate that nitrate-dependent metabolism promotes adaptation of S. Typhimurium to bile.

Keywords: RNA‐seq; Salmonella; bile; metabolism; nitrate; stress.

MeSH terms

  • Bacterial Proteins / genetics
  • Bacterial Proteins / metabolism
  • Bile / metabolism
  • Gene Expression Profiling
  • Gene Expression Regulation, Bacterial
  • Nitrates* / metabolism
  • Reactive Oxygen Species / metabolism
  • Salmonella typhimurium* / genetics
  • Salmonella typhimurium* / metabolism
  • Stress, Physiological / genetics
  • Transcriptome

Substances

  • Nitrates
  • Bacterial Proteins
  • Reactive Oxygen Species