1 Washington University in St. Louis School of Medicine (LZ), St. Louis, Missouri; Department of Pathology and Immunology (KR), Washington University School of Medicine, St. Louis, Missouri; Mercy Clinic Eye Specialists (HC), St. Louis, Missouri; and John F. Hardesty, MD Department of Ophthalmology and Visual Sciences and Department of Neurology (GVS, LS), Washington University School of Medicine, St. Louis, Missouri.