Ketogenic diet administration later in life improves memory by modifying the synaptic cortical proteome via the PKA signaling pathway in aging mice

Cell Rep Med. 2024 Jun 18;5(6):101593. doi: 10.1016/j.xcrm.2024.101593. Epub 2024 Jun 5.

Abstract

Aging compromises brain function leading to cognitive decline. A cyclic ketogenic diet (KD) improves memory in aged mice after long-term administration; however, short-term effects later in life and the molecular mechanisms that govern such changes remain unclear. Here, we explore the impact of a short-term KD treatment starting at elderly stage on brain function of aged mice. Behavioral testing and long-term potentiation (LTP) recordings reveal that KD improves working memory and hippocampal LTP. Furthermore, the synaptosome proteome of aged mice fed a KD long-term evidence changes predominantly at the presynaptic compartment associated to the protein kinase A (PKA) signaling pathway. These findings were corroborated in vivo by western blot analysis, with high BDNF abundance and PKA substrate phosphorylation. Overall, we show that a KD modifies brain function even when it is administered later in life and recapitulates molecular features of long-term administration, including the PKA signaling pathway, thus promoting synaptic plasticity at advanced age.

Keywords: BDNF; LTP; PKA; aging; brain-derived neurotrophic factor; ketogenic diet; long-term potentiation; protein kinase A; proteomics; β-hydroxybutyrate.

MeSH terms

  • Aging* / metabolism
  • Aging* / physiology
  • Animals
  • Brain-Derived Neurotrophic Factor / metabolism
  • Cyclic AMP-Dependent Protein Kinases* / metabolism
  • Diet, Ketogenic* / methods
  • Hippocampus / metabolism
  • Long-Term Potentiation* / physiology
  • Male
  • Memory* / physiology
  • Mice
  • Mice, Inbred C57BL
  • Neuronal Plasticity / physiology
  • Phosphorylation
  • Proteome* / metabolism
  • Signal Transduction*
  • Synapses / metabolism

Substances

  • Cyclic AMP-Dependent Protein Kinases
  • Proteome
  • Brain-Derived Neurotrophic Factor