Treponema pallidum Periplasmic and Membrane Proteins Are Recognized by Circulating and Skin CD4+ T Cells

J Infect Dis. 2024 Aug 16;230(2):281-292. doi: 10.1093/infdis/jiae245.

Abstract

Background: Histologic and serologic studies suggest the induction of local and systemic Treponema pallidum-specific CD4+ T-cell responses to T. pallidum infection. We hypothesized that T. pallidum-specific CD4+ T cells are detectable in blood and in the skin rash of secondary syphilis and persist in both compartments after treatment.

Methods: Peripheral blood mononuclear cells collected from 67 participants were screened by interferon-γ (IFN-γ) ELISPOT response to T. pallidum sonicate. T. pallidum-reactive T-cell lines from blood and skin were probed for responses to 89 recombinant T. pallidum antigens. Peptide epitopes and HLA class II restriction were defined for selected antigens.

Results: We detected CD4+ T-cell responses to T. pallidum sonicate ex vivo. Using T. pallidum-reactive T-cell lines we observed recognition of 14 discrete proteins, 13 of which localize to bacterial membranes or the periplasmic space. After therapy, T. pallidum-specific T cells persisted for at least 6 months in skin and 10 years in blood.

Conclusions: T. pallidum infection elicits an antigen-specific CD4+ T-cell response in blood and skin. T. pallidum-specific CD4+ T cells persist as memory in both compartments long after curative therapy. The T. pallidum antigenic targets we identified may be high-priority vaccine candidates.

Keywords: CD4+ T cell; interferon-γ; secondary syphilis; syphilis; tissue-resident memory T cell; vaccine.

MeSH terms

  • Adult
  • Antigens, Bacterial / immunology
  • Bacterial Proteins / immunology
  • CD4-Positive T-Lymphocytes* / immunology
  • Enzyme-Linked Immunospot Assay
  • Female
  • Humans
  • Interferon-gamma / metabolism
  • Leukocytes, Mononuclear / immunology
  • Male
  • Membrane Proteins / immunology
  • Middle Aged
  • Skin* / immunology
  • Skin* / microbiology
  • Syphilis* / immunology
  • Treponema pallidum* / immunology
  • Young Adult

Substances

  • Membrane Proteins
  • Antigens, Bacterial
  • Interferon-gamma
  • Bacterial Proteins

Supplementary concepts

  • Syphilis, secondary