SARS-CoV-2 nucleocapsid protein promotes self-deacetylation by inducing HDAC6 to facilitate viral replication

Virol J. 2024 Aug 12;21(1):186. doi: 10.1186/s12985-024-02460-5.

Abstract

Background: The global outbreak of COVID-19 caused by the SARS-CoV-2 has led to millions of deaths. This unanticipated emergency has prompted virologists across the globe to delve deeper into the intricate dynamicity of the host-virus interface with an aim to identify antiviral targets and elucidate host and viral determinants of severe disease.

Aim: The present study was undertaken to analyse the role of histone deacetylase 6 (HDAC6) in regulating SARS-CoV-2 infection.

Results: Gradual increase in HDAC6 expression was observed in different SARS-CoV-2-permissive cell lines following SARS-CoV-2 infection. The SARS-CoV-2 nucleocapsid protein (N protein) was identified as the primary viral factor responsible for upregulating HDAC6 expression. Downregulation of HDAC6 using shRNA or a specific inhibitor tubacin resulted in reduced viral replication suggesting proviral role of its deacetylase activity. Further investigations uncovered the interaction of HDAC6 with stress granule protein G3BP1 and N protein during infection. HDAC6-mediated deacetylation of SARS-CoV-2 N protein was found to be crucial for its association with G3BP1.

Conclusion: This study provides valuable insights into the molecular mechanisms underlying the disruption of cytoplasmic stress granules during SARS-CoV-2 infection and highlights the significance of HDAC6 in the process.

Keywords: Deacetylation; G3BP1; HDAC6; SARS-CoV-2; Stress granules (SGs).

MeSH terms

  • Acetylation
  • Animals
  • COVID-19* / metabolism
  • COVID-19* / virology
  • Cell Line
  • Chlorocebus aethiops
  • Coronavirus Nucleocapsid Proteins* / genetics
  • Coronavirus Nucleocapsid Proteins* / metabolism
  • DNA Helicases
  • Histone Deacetylase 6* / genetics
  • Histone Deacetylase 6* / metabolism
  • Host-Pathogen Interactions
  • Humans
  • Phosphoproteins / genetics
  • Phosphoproteins / metabolism
  • Poly-ADP-Ribose Binding Proteins
  • RNA Helicases
  • RNA Recognition Motif Proteins / metabolism
  • SARS-CoV-2* / physiology
  • Vero Cells
  • Virus Replication*

Substances

  • Histone Deacetylase 6
  • HDAC6 protein, human
  • Coronavirus Nucleocapsid Proteins
  • G3BP1 protein, human
  • RNA Recognition Motif Proteins
  • nucleocapsid phosphoprotein, SARS-CoV-2
  • Phosphoproteins
  • Poly-ADP-Ribose Binding Proteins
  • DNA Helicases
  • RNA Helicases