1 Laboratory of Plant Physiology, Plant Sciences Group, Wageningen University and Research,Wageningen 6708 PB, the Netherlands.
2 Biological and Environmental Science and Engineering Division, King Abdullah University of Science and Technology (KAUST), Thuwal 23955-6900, Kingdom of Saudi Arabia.
3 School of Biological Sciences, College of Natural Science, Seoul National University, Seoul 08826, Korea.
4 Faculty of Biology, Technion-Israel Institute of Technology, Haifa 3200003, Israel.