Abstract
For a brief period during fetal lymph node organogenesis in mice, lymph node postcapillary high endothelial venules surprisingly express the Peyer's patch addressin MAdCAM-1. This expression allows initial seeding of this incipient structure by two unusual lymphocyte populations selectively expressing the Peyer's patch homing receptor integrin alpha4beta7: CD4+CD3- oligolineage progenitors and TCR gammadelta+ T cells. We show here that CD4+CD3- cells are lineage-restricted progenitors that express surface lymphotoxin-beta (LTbeta) and the chemokine receptor BLR1 and that can become natural killer cells, dendritic antigen-presenting cells, and follicular cells of unknown outcome, but these cells do not become T or B lymphocytes. Since the necessity of lymphotoxin in lymphoid organ development has been shown, we propose that the novel subset of CD4+CD3-LTbeta+ fetal cells is instrumental in the development of lymphoid tissue architecture.
Publication types
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Research Support, Non-U.S. Gov't
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Research Support, U.S. Gov't, P.H.S.
MeSH terms
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Animals
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Animals, Newborn
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Antigen-Presenting Cells / cytology*
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B-Lymphocytes / cytology
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CD3 Complex / metabolism*
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CD4-Positive T-Lymphocytes / cytology*
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Cell Adhesion Molecules
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Cytotoxicity, Immunologic
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Fluorescent Antibody Technique, Indirect
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GTP-Binding Proteins / genetics
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Gene Expression
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Histocompatibility Antigens Class II / metabolism
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Immunity, Cellular
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Immunoglobulins / metabolism
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Integrins / metabolism
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Interleukin-2 / pharmacology
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Killer Cells, Natural / cytology*
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Leukocyte Common Antigens / analysis
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Leukopoiesis*
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Lymph Nodes / cytology
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Lymph Nodes / embryology*
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Lymphocyte Subsets / cytology*
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Lymphotoxin-alpha / metabolism*
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Lymphotoxin-beta
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Membrane Proteins / metabolism*
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Mice
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Mice, Inbred AKR
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Mice, Inbred BALB C
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Mice, Inbred C57BL
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Mucoproteins / metabolism
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RNA, Messenger / genetics
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Receptors, CXCR5
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Receptors, Chemokine
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Receptors, Cytokine / genetics
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Spleen / embryology
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Spleen / immunology
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T-Lymphocytes / cytology
Substances
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CD3 Complex
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CXCR5 protein, mouse
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Cell Adhesion Molecules
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Histocompatibility Antigens Class II
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Immunoglobulins
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Integrins
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Interleukin-2
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LTB protein, human
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Ltb protein, mouse
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Lymphotoxin-alpha
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Lymphotoxin-beta
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Madcam1 protein, mouse
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Membrane Proteins
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Mucoproteins
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RNA, Messenger
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Receptors, CXCR5
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Receptors, Chemokine
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Receptors, Cytokine
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integrin alpha4beta7
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Leukocyte Common Antigens
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GTP-Binding Proteins