Abstract
Insulin-like growth factor binding proteins (IGFBPs) are important modulators of IGF actions. IGFBP-3 and IGFBP-5 can bind to the extracellular matrix of a number of cell types. We now describe a new posttranslational structural modification of IGFBP-3 and IGFBP-5, which could play a role in determining their localization. We incubated radioiodinated forms of all six IGFBPs in the presence of a redox buffer consisting of 10 mM reduced glutathione and 0.2 mM oxidized glutathione. Under these conditions IGFBP-3 and IGFBP-5, but not the other IGFBPs, formed high molecular weight disulfide-linked multimers. Heparin and a peptide encompassing the high-affinity heparin-binding site in the C-terminal portion of IGFBP-3 were capable of blocking the multimerization of IGFBP-3. IGFBP-3, but not IGFBP-1, was shown to be able to self-associate non-covalently, which could be a requisite first step in the formation of covalent multimers. The self-association of IGFBP-3 required the high-affinity heparin-binding site in the C-terminal portion of the molecule.
MeSH terms
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Amino Acid Sequence
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Binding Sites
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Chromatography, Gel
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Electrophoresis, Polyacrylamide Gel
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Glutathione / pharmacology
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Glutathione Disulfide / pharmacology
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Heparin / pharmacology
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Humans
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Insulin-Like Growth Factor Binding Protein 3 / chemistry*
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Insulin-Like Growth Factor Binding Protein 3 / metabolism
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Insulin-Like Growth Factor Binding Protein 5 / chemistry*
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Insulin-Like Growth Factor Binding Protein 5 / metabolism
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Insulin-Like Growth Factor I / metabolism
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Iodine Radioisotopes / analysis
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Iodine Radioisotopes / metabolism
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Macromolecular Substances
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Molecular Sequence Data
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Molecular Weight
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Oxidation-Reduction
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Peptide Fragments / pharmacology
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Protein Conformation*
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Recombinant Proteins / chemistry
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Recombinant Proteins / metabolism
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Sodium Dodecyl Sulfate / pharmacology*
Substances
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Insulin-Like Growth Factor Binding Protein 3
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Insulin-Like Growth Factor Binding Protein 5
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Iodine Radioisotopes
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Macromolecular Substances
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Peptide Fragments
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Recombinant Proteins
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Sodium Dodecyl Sulfate
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Insulin-Like Growth Factor I
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Heparin
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Glutathione
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Glutathione Disulfide