Abstract
Signaling pathways that stabilize interleukin-2 (IL-2) messenger RNA (mRNA) in activated T cells were examined. IL-2 mRNA contains at least two cis elements that mediated its stabilization in response to different signals, including activation of c-Jun amino-terminal kinase (JNK). This response was mediated through a cis element encompassing the 5' untranslated region (UTR) and the beginning of the coding region. IL-2 transcripts lacking this 5' element no longer responded to JNK activation but were still responsive to other signals generated during T cell activation, which were probably sensed through the 3' UTR. Thus, multiple elements within IL-2 mRNA modulate its stability in a combinatorial manner, and the JNK pathway controls turnover as well as synthesis of IL-2 mRNA.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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CD28 Antigens / immunology
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CD3 Complex / immunology
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Calcimycin / pharmacology
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Calcium-Calmodulin-Dependent Protein Kinases / antagonists & inhibitors
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Calcium-Calmodulin-Dependent Protein Kinases / metabolism*
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Cyclosporine / pharmacology
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Enzyme Activation
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Enzyme Inhibitors / pharmacology
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Gene Expression Regulation*
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Humans
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Imidazoles / pharmacology
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Interleukin-2 / genetics*
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JNK Mitogen-Activated Protein Kinases
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Jurkat Cells
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Lymphocyte Activation
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MAP Kinase Kinase 1
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MAP Kinase Kinase 7
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Mitogen-Activated Protein Kinase Kinases*
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Mitogen-Activated Protein Kinases*
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Protein Kinases / metabolism
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Protein Serine-Threonine Kinases / metabolism
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Protein-Tyrosine Kinases / metabolism
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Pyridines / pharmacology
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RNA, Messenger / chemistry
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RNA, Messenger / genetics
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RNA, Messenger / metabolism*
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Signal Transduction
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T-Lymphocytes / immunology
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T-Lymphocytes / metabolism*
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Tetradecanoylphorbol Acetate / pharmacology
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Transgenes
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p38 Mitogen-Activated Protein Kinases
Substances
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CD28 Antigens
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CD3 Complex
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Enzyme Inhibitors
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Imidazoles
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Interleukin-2
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Pyridines
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RNA, Messenger
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Calcimycin
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Cyclosporine
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Protein Kinases
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Protein-Tyrosine Kinases
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Protein Serine-Threonine Kinases
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Calcium-Calmodulin-Dependent Protein Kinases
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JNK Mitogen-Activated Protein Kinases
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Mitogen-Activated Protein Kinases
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p38 Mitogen-Activated Protein Kinases
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MAP Kinase Kinase 1
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MAP Kinase Kinase 7
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MAP2K1 protein, human
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MAP2K7 protein, human
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Mitogen-Activated Protein Kinase Kinases
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Tetradecanoylphorbol Acetate
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4-(4-fluorophenyl)-2-(4-hydroxyphenyl)-5-(4-pyridyl)imidazole