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H4N8 subtype avian influenza virus isolated from shorebirds contains a unique PB1 gene and causes severe respiratory disease in mice.
Bui VN, Ogawa H, Xininigen, Karibe K, Matsuo K, Awad SS, Minoungou GL, Yoden S, Haneda H, Ngo LH, Tamaki S, Yamamoto Y, Nakamura K, Saito K, Watanabe Y, Runstadler J, Huettmann F, Happ GM, Imai K. Bui VN, et al. Virology. 2012 Feb 5;423(1):77-88. doi: 10.1016/j.virol.2011.11.019. Epub 2011 Dec 20. Virology. 2012. PMID: 22192630 Free PMC article.