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'''''Manihot esculenta''''', [[common name|commonly called]] '''cassava'''
Cassava is the third-largest source of carbohydrates in food in the tropics, after [[rice]] and [[maize]], making it is an important [[staple food|staple]]; more than 500 million people depend on it. It offers the advantage of being exceptionally [[drought-tolerant]], and able to grow productively on poor soil. The largest producer is Nigeria, while Thailand is the largest exporter of cassava starch.
Cassava is
==Description==
The harvested part of a cassava plant is the root. This is long and tapered, with an easily
<gallery class=center mode=nolines widths=180 heights=180>
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[[File:Albert_Eckhout_-_Mandioca.jpg|thumb|17th-century painting by [[Albert Eckhout]] in [[Dutch Brazil]]]]
Wild populations of ''M. esculenta'' subspecies ''flabellifolia'', shown to be the progenitor of domesticated cassava, are centered in west-central Brazil, where it was likely first domesticated no more than 10,000 years ago.<ref>{{Cite journal|last1=Olsen|first1=K. M. |last2=Schaal |first2=B. A. |title=Evidence on the origin of cassava: phylogeography of ''Manihot esculenta'' |journal=[[Proceedings of the National Academy of Sciences of the United States of America]]|volume=96 |issue=10 |pages=5586–5591 |year=1999 |pmid=10318928 |pmc=21904 |bibcode=1999PNAS...96.5586O |doi=10.1073/pnas.96.10.5586 |doi-access=free }}</ref> Forms of the modern domesticated species can also be found growing in the wild in the south of Brazil. By 4600 BC, cassava pollen appears in the [[Gulf of Mexico]] lowlands, at the [[San Andrés (Mesoamerican site)|San Andrés]] archaeological site.<ref>{{cite journal |doi=10.1126/science.292.5520.1370 |pmid=11359011 |bibcode=2001Sci...292.1370P |title=Origin and Environmental Setting of Ancient Agriculture in the Lowlands of Mesoamerica |journal=[[Science (journal)|Science]] |volume=292 |issue=5520 |pages=1370–1373 |last1=Pope |first1=Kevin O. |last2=Pohl |first2=Mary E. D. |last3=Jones |first3=John G. |last4=Lentz |first4=David L. |last5=von Nagy |first5=Christopher |last6=Vega |first6=Francisco J. |last7=Quitmyer |first7=Irvy R. |year=2001}}</ref> The oldest direct evidence of cassava cultivation comes from a 1,400-year-old [[Maya civilization|Maya]] site, [[Joya de Cerén]], in [[El Salvador]].<ref>{{cite news |last=Carroll |first=Rory |title=CU team discovers Mayan crop system |newspaper=[[The Guardian]] |date=23 August 2007 |url=https://www.theguardian.com/science/2007/aug/23/1 |access-date=31 July 2019 |archive-date=31 July 2019 |archive-url=https://web.archive.org/web/20190731171801/https://www.theguardian.com/science/2007/aug/23/1 |url-status=live }}</ref>
[[Spaniards]] in their early occupation of Caribbean islands did not want to eat cassava or maize, which they considered insubstantial, dangerous, and not nutritious. They much preferred foods from Spain, specifically wheat bread, olive oil, red wine, and meat, and considered maize and cassava damaging to Europeans.<ref>Earle, Rebecca (2012) ''The Body of the Conquistador: Food, Race, and the Colonial Experience in Spanish America, 1492–1700''. New York: Cambridge University Press. pp. 54–57, 151. {{ISBN|978-1107693296}}.</ref> The cultivation and consumption of cassava were nonetheless continued in both Portuguese and Spanish America. Mass production of cassava bread became the first Cuban industry established by the Spanish.<ref>{{cite book|author=Long, Janet |url=https://books.google.com/books?id=Rn9LZ2XrIWgC&pg=PA75 |title=Conquest and food: consequences of the encounter of two worlds; page 75|publisher=UNAM|date=2003 |isbn=978-9703208524 |access-date=24 August 2020 |archive-date=20 April 2023|archive-url=https://web.archive.org/web/20230420125914/https://books.google.com/books?id=Rn9LZ2XrIWgC&pg=PA75 |url-status=live}}</ref> Ships departing to Europe from Cuban ports such as [[Havana]], [[Santiago de Cuba|Santiago]], [[Bayamo]], and [[Baracoa]] carried goods to Spain, but sailors needed to be provisioned for the voyage. The Spanish also needed to replenish their boats with dried meat, water, fruit, and large amounts of cassava bread.<ref>{{cite web|url=http://www.sjsu.edu/faculty/watkins/havana.htm |title=The Economic History of Havana, Cuba: A City So Beautiful and Important It Was Once Worth More Than All of Florida |author=Watkins, Thayer|publisher=San José State University, Department of Economics|date=2006|access-date=20 August 2015 |archive-url=https://web.archive.org/web/20160502042108/http://www.sjsu.edu/faculty/watkins/havana.htm |archive-date=2 May 2016}}</ref> Sailors complained that it caused them digestive problems.<ref>{{cite journal |author=Super, John C. |title=Spanish Diet in the Atlantic Crossing|journal=[[Terrae Incognitae]] |volume=16 |year=1984 |pages=60–63 |doi=10.1179/008228884791016718}}</ref>
There is a legend that cassava was introduced in 1880–1885 CE to the South Indian state of [[Kerala]] by the King of [[Travancore]], Vishakham Thirunal Maharaja, after a great famine hit the kingdom, as a substitute for rice.<ref name=Saraswathy2019>{{cite web |date=June 27, 2019 |author=Saraswathy Nagarajan |url=https://www.thehindu.com/life-and-style/food/how-tapioca-came-to-travancore/article28181288.ece |title=How tapioca came to Travancore |work=The Hindu |archive-url=https://web.archive.org/web/20200727082403/https://www.thehindu.com/life-and-style/food/how-tapioca-came-to-travancore/article28181288.ece |archive-date=27 July 2020 |url-status=live}}</ref> However, there are documented cases of cassava cultivation in parts of the state before the time of Vishakham Thirunal Maharaja.<ref>{{Cite book |last1=Ainslie |first1=Whitelaw |url=http://archive.org/details/b28037340 |title=Materia medica of Hindoostan, and artisan's and agriculturalist's nomenclature |last2=Halford|first2=Henry|date=1813|publisher=Government Press |location=Madras State}}</ref> Cassava is called kappa or maricheeni in [[Malayalam]], and [[tapioca]] in Indian English usage.<ref>{{cite web |title=Kappa for all seasons - many avatars of the magic starch root... |url=https://www.onmanorama.com/food/features/2018/01/30/kappa-for-all-seasons.html |website=Onmanorama |location=[[Kerala]], India |access-date=11 May 2024 |date=1 February 2018}}</ref>
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</gallery>
=== Pests ===
{{further|List of cassava diseases}}
[[File:Thailand 19 lo (4039130033).jpg|thumb|An agronomist examines a diseased cassava crop in Thailand.]]
Many species of insects contribute to serious losses, between 19% and 30%, of dried cassava in storage.<ref name="jis">{{cite journal |last1=Osipitan |first1=A. A. |last2=Sangowusi |first2=V. T. |last3=Lawal |first3=O. I. |last4=Popoola |first4=K. O. |year=2015 |title=Correlation of Chemical Compositions of Cassava Varieties to Their Resistance to ''Prostephanus truncatus'' Horn (Coleoptera: Bostrichidae) |journal=[[Journal of Insect Science (Entomological Society of America)|Journal of Insect Science]]|volume=15 |issue=1 |page=13 |doi=10.1093/jisesa/ieu173 |pmc=4535132 |pmid=25700536}}</ref> In [[Africa]], a previous issue was the cassava mealybug (''[[Phenacoccus manihoti]]'') and cassava green mite (''[[Mononychellus tanajoa]]''). These pests can cause up to 80 percent crop loss, which is extremely detrimental to the production of [[subsistence economy|subsistence]] farmers. These pests were rampant in the 1970s and 1980s but were brought under control following the establishment of the Biological Control Centre for Africa of the [[International Institute of Tropical Agriculture]] (IITA) under the leadership of [[Hans Rudolf Herren]].<ref>{{cite web |title=1995: Herren |url=http://www.worldfoodprize.org/en/laureates/19871999_laureates/1995_herren/ |archive-url=https://web.archive.org/web/20150711200345/http://www.worldfoodprize.org/en/laureates/19871999_laureates/1995_herren/ |archive-date=11 July 2015 |access-date=29 May 2015 |publisher=The World Food Prize Foundation}}</ref> The Centre investigated [[biological control]] for cassava pests; two [[South America]]n natural enemies ''Anagyrus lopezi'' (a [[parasitoid]] wasp) and ''[[Typhlodromalus]] aripo'' (a predatory mite) were found to effectively control the cassava mealybug and the cassava green mite, respectively.<ref>{{cite web |title=1995: Herren |url=http://www.worldfoodprize.org/en/laureates/19871999_laureates/1995_herren/ |archive-url=https://web.archive.org/web/20150711200345/http://www.worldfoodprize.org/en/laureates/19871999_laureates/1995_herren/ |archive-date=11 July 2015 |access-date=29 May 2015 |publisher=The World Food Prize Foundation}}</ref>▼
Cassava is subject to pests from multiple taxonomic groups, including viruses, bacteria, fungi, nematodes, and insects. All cause reductions in yield, and some cause serious losses of crops.<ref name="Alvarez 2012"/>
[[Xanthomonas axonopodis pv. manihotis|''Xanthomonas axonopodis'' pv. ''manihotis'']] causes [[bacterial blight of cassava]]. This disease originated in South America and has followed cassava around the world.<ref name="Lozano-1986">{{Cite journal |last=Lozano |first=J. Carlos |date=September 1986 |title=Cassava bacterial blight: a manageable disease |url=http://www.apsnet.org/publications/PlantDisease/BackIssues/Documents/1986Articles/PlantDisease70n12_1089.PDF |journal=[[Plant Disease (journal)|Plant Disease]] |publisher=[[American Phytopathological Society]] (APS) |volume=70 |issue=12 |pages=1089–1093 |doi=10.1094/PD-70-1089 |access-date=14 January 2023 |archive-date=14 January 2023 |archive-url=https://web.archive.org/web/20230114224139/https://www.apsnet.org/publications/PlantDisease/BackIssues/Documents/1986Articles/PlantDisease70n12_1089.PDF |url-status=live }}</ref> Bacterial blight has been responsible for near catastrophic losses and famine in past decades, and its mitigation requires active management practices.<ref name="Lozano-1986" /> Several other bacteria attack cassava, including the related ''[[Xanthomonas campestris]]'' pv. ''cassavae'', which causes bacterial angular leaf spot.<ref name="Zárate‐Chaves 2021">{{cite journal |last1=Zárate-Chaves |first1=Carlos A. |last2=Gómez de la Cruz |first2=Diana |last3=Verdier |first3=Valérie |last4=López |first4=Camilo E. |last5=Bernal |first5=Adriana |last6=Szurek |first6=Boris |title=Cassava diseases caused by Xanthomonas phaseoli pv. manihotis and Xanthomonas cassavae |journal=Molecular Plant Pathology |volume=22 |issue=12 |date=2021 |issn=1464-6722 |pmid=34227737 |pmc=8578842 |doi=10.1111/mpp.13094 |pages=1520–1537}}</ref>▼
; Viruses
Several viruses cause enough damage to cassava crops to be of economic importance. The [[African cassava mosaic virus]] causes the leaves of the cassava plant to wither, limiting the growth of the root.<ref>{{cite web |title=Cassava
; Bacteria
A wide range of plant parasitic nematodes have been reported associated with cassava worldwide. These include ''[[Pratylenchus brachyurus]]'', ''[[Rotylenchulus reniformis]]'', ''[[Helicotylenchus]]'' spp., ''[[Scutellonema]]'' spp. and ''[[Root-knot nematode|Meloidogyne]]'' spp., of which ''[[Meloidogyne incognita]]'' and ''[[Meloidogyne javanica]]'' are the most widely reported and economically important.<ref>{{cite journal |author1=Mc Sorley, R. |author2=Ohair, S. K. |author3=Parrado, J. L. |date=1983 |title=Nematodes of Cassava, ''Manihot esculenta'' Crantz |url=http://journals.fcla.edu/nematropica/article/view/63820/61488 |journal=Nematropica|volume=13 |pages=261–287 |archive-url=https://web.archive.org/web/20160603205439/http://journals.fcla.edu/nematropica/article/view/63820/61488 |archive-date=3 June 2016 |access-date=4 May 2016}}</ref> ''Meloidogyne'' spp. feeding produces physically damaging galls with eggs inside them. Galls later merge as the females grow and enlarge, and they interfere with water and nutrient supply.<ref name="Gapasin">{{cite journal |author1=Gapasin, R. M. |date=1980 |title=Reaction of golden yellow cassava to ''Meloidogyne'' spp. Inoculation |journal=Annals of Tropical Research|volume=2 |pages=49–53}}</ref> Cassava roots become tough with age and restrict the movement of the juveniles and the egg release. It is therefore possible that extensive galling can be observed even at low densities following infection.<ref name="Coyne">{{cite journal |author1=Coyne, D. L. |date=1994 |title=Nematode pests of cassava |journal=African Crop Science Journal|volume=2 |issue=4 |pages=355–359 |url=https://www.ajol.info/index.php/acsj/article/view/135776 |access-date=22 September 2018 |archive-date=22 September 2018 |url-status=live |archive-url=https://web.archive.org/web/20180922101847/https://www.ajol.info/index.php/acsj/article/view/135776}}</ref> Other pests and diseases can gain entry through the physical damage caused by gall formation, leading to rots. They have not been shown to cause direct damage to the enlarged storage roots, but plants can have reduced height if there was loss of enlarged root weight.<ref>{{cite journal |author1=Caveness, F. E. |date=1982 |title=Root-knot nematodes as parasites of cassava |journal=IITA Research Briefs|volume=3 |issue=2 |pages=2–3}}</ref>▼
▲Among the most serious bacterial pests is [[Xanthomonas axonopodis pv. manihotis|''Xanthomonas axonopodis'' pv. ''manihotis'']], which causes [[bacterial blight of cassava]]. This disease originated in South America and has followed cassava around the world.<ref name="Lozano-1986">{{Cite journal |last=Lozano |first=J. Carlos |date=September 1986 |title=Cassava bacterial blight: a manageable disease |url=http://www.apsnet.org/publications/PlantDisease/BackIssues/Documents/1986Articles/PlantDisease70n12_1089.PDF |journal=[[Plant Disease (journal)|Plant Disease]] |publisher=[[American Phytopathological Society]] (APS) |volume=70 |issue=12 |pages=1089–1093 |doi=10.1094/PD-70-1089 |access-date=14 January 2023 |archive-date=14 January 2023 |archive-url=https://web.archive.org/web/20230114224139/https://www.apsnet.org/publications/PlantDisease/BackIssues/Documents/1986Articles/PlantDisease70n12_1089.PDF |url-status=live }}</ref> Bacterial blight has been responsible for near catastrophic losses and famine in past decades, and its mitigation requires active management practices.<ref name="Lozano-1986" /> Several other bacteria attack cassava, including the related ''[[Xanthomonas campestris]]'' pv. ''cassavae'', which causes bacterial angular leaf spot.<ref name="Zárate‐Chaves 2021">{{cite journal |last1=Zárate-Chaves |first1=Carlos A. |last2=Gómez de la Cruz |first2=Diana |last3=Verdier |first3=Valérie |last4=López |first4=Camilo E. |last5=Bernal |first5=Adriana |last6=Szurek |first6=Boris |title=Cassava diseases caused by Xanthomonas phaseoli pv. manihotis and Xanthomonas cassavae |journal=Molecular Plant Pathology |volume=22 |issue=12 |date=2021 |issn=1464-6722 |pmid=34227737 |pmc=8578842 |doi=10.1111/mpp.13094 |pages=1520–1537}}</ref>
; Fungi
The use of nematicides has been found to result in lower numbers of galls per feeder root compared to a control, coupled with a lower number of rots in the storage roots.<ref>{{cite journal |author1=Coyne D. L. |author2=Kagoda F. |author3=Wambugu E. |author4=Ragama P. |date=2006 |title=Response of cassava to nematicide application and plant-parasitic nematode infection in East Africa, with emphasis on root-knot nematode |journal=International Journal of Pest Management |volume=52 |issue=3 |pages=215–223 |doi=10.1080/09670870600722959 |s2cid=84771539}}</ref> The organophosphorus nematicide femaniphos, when used, did not affect crop growth and yield parameter variables measured at harvest. Nematicide use in cassava is not terribly effective at increasing harvested yield, but lower infestation at harvest and lower subsequent storage loss provide a higher effective yield. The use of tolerant and resistant cultivars is the most practical management method in most locales.<ref name="Coyne-et-al-2018">{{cite journal |last1=Coyne |first1=Danny L. |last2=Cortada |first2=Laura |last3=Dalzell |first3=Johnathan J. |last4=Claudius-Cole |first4=Abiodun O. |last5=Haukeland |first5=Solveig |last6=Luambano |first6=Nessie |last7=Talwana |first7=Herbert |date=2018-08-25 |title=Plant-Parasitic Nematodes and Food Security in Sub-Saharan Africa |journal=[[Annual Review of Phytopathology]] |publisher=[[Annual Reviews (publisher)|Annual Reviews]] |volume=56 |issue=1 |pages=381–403 |doi=10.1146/annurev-phyto-080417-045833 |issn=0066-4286 |pmc=7340484 |pmid=29958072 |s2cid=49615468}}</ref><ref name="Coyne"/><ref name="TodayNg-new-varieties">{{cite web |url=http://www.today.ng/news/nigeria/nigeria-introduce-cassava-varieties-334463 |title=Nigeria to introduce new cassava varieties |website=Todayng |date=2020-12-21 |first=Chidinma |last=Uchechukwumgemezu |access-date=21 December 2020 |archive-date=21 December 2020 |archive-url=https://web.archive.org/web/20201221194624/https://www.today.ng/news/nigeria/nigeria-introduce-cassava-varieties-334463 |url-status=live}}</ref>▼
{{anchor|Rust of cassava}}
Several fungi bring about significant crop losses, one of the most serious being cassava root rot; the [[pathogen]]s involved are species of ''[[Phytophthora]]'', the genus which causes potato blight. Cassava root rot can result in losses of as much as 80 percent of the crop.<ref name="Alvarez 2012">{{cite book|last1=Alvarez |first1=Elizabeth |author2=Llano, Germán Alberto |author3=Mejía, Juan Fernando |chapter=Cassava diseases in Latin America, Africa and Asia |title=The Cassava Handbook |page=258 |year=2012 |url=https://www.academia.edu/download/98098352/The_Cassava_Handbook_2011.pdf#page=266}}<!--Scholar: cassava fungal diseases--></ref>
Superelongation disease, caused by ''[[Elsinoë brasiliensis]]'', can cause losses of over 80 percent of young cassava in Latin America when temperature and rainfall are high.<ref name="Alvarez 2012"/>
; Nematodes
▲This crop suffers from a [[rust (fungus)|rust]], {{Visible anchor|rust of cassava|Uromyces manihotis}}, caused by ''Uromyces manihotis''.<ref name="Rust">{{Cite web |url=https://www.cabi.org/isc/datasheet/55848 |title=''Uromyces manihotis'' (rust of cassava) |year=2019 |website=[[Invasive Species Compendium]] (ISC) |publisher=[[Centre for Agriculture and Bioscience International|CABI]] (Centre for Agriculture and Bioscience International) |access-date=2022-10-27 |archive-date=9 November 2022 |archive-url=https://web.archive.org/web/20221109225057/https://www.cabi.org/isc/datasheet/55848 |url-status=live }}</ref>
▲Nematode pests of cassava are thought to cause harms ranging from negligible to seriously damaging,<ref>{{cite journal |author1=Coyne, D. L. |author2=Talwana, L. A. H. |date=2000 |title=Reaction of cassava cultivars to root-knot nematode (''Meloidogyne'' spp.) in pot experiments and farmer-managed field trials in Uganda |journal=International Journal of Nematology|volume=10 |pages=153–158 |url=https://www.cabdirect.org/cabdirect/abstract/20013117434 |access-date=22 September 2018}}</ref><ref>{{cite journal |author1=Makumbi-Kidza, N. N. |author2=Speijer |author3=Sikora R. A. |date=2000 |title=Effects of ''Meloidogyne incognita'' on growth and storage-root formation of cassava (''Manihot esculenta'') |journal=Journal of Nematology|volume=32 |issue=4S |pages=475–477 |pmc=2620481 |pmid=19270997}}</ref><ref name="Gapasin"/> making the choice of management methods difficult.<ref name="Coyne"/> A wide range of plant parasitic nematodes have been reported associated with cassava worldwide. These include ''[[Pratylenchus brachyurus]]'', ''[[Rotylenchulus reniformis]]'', ''[[Helicotylenchus]]'' spp., ''[[Scutellonema]]'' spp. and ''[[Root-knot nematode|Meloidogyne]]'' spp., of which ''[[Meloidogyne incognita]]'' and ''[[Meloidogyne javanica]]'' are the most widely reported and economically important.<ref>{{cite journal |author1=Mc Sorley, R. |author2=Ohair, S. K. |author3=Parrado, J. L. |date=1983 |title=Nematodes of Cassava, ''Manihot esculenta'' Crantz |url=http://journals.fcla.edu/nematropica/article/view/63820/61488 |journal=Nematropica|volume=13 |pages=261–287 |archive-url=https://web.archive.org/web/20160603205439/http://journals.fcla.edu/nematropica/article/view/63820/61488 |archive-date=3 June 2016 |access-date=4 May 2016}}</ref> ''Meloidogyne'' spp. feeding produces physically damaging galls with eggs inside them. Galls later merge as the females grow and enlarge, and they interfere with water and nutrient supply.<ref name="Gapasin">{{cite journal |author1=Gapasin, R. M. |date=1980 |title=Reaction of golden yellow cassava to ''Meloidogyne'' spp. Inoculation |journal=Annals of Tropical Research|volume=2 |pages=49–53}}</ref> Cassava roots become tough with age and restrict the movement of the juveniles and the egg release. It is therefore possible that extensive galling can be observed even at low densities following infection.<ref name="Coyne">{{cite journal |author1=Coyne, D. L. |date=1994 |title=Nematode pests of cassava |journal=African Crop Science Journal|volume=2 |issue=4 |pages=355–359 |url=https://www.ajol.info/index.php/acsj/article/view/135776 |access-date=22 September 2018 |archive-date=22 September 2018 |url-status=live |archive-url=https://web.archive.org/web/20180922101847/https://www.ajol.info/index.php/acsj/article/view/135776}}</ref> Other pests and diseases can gain entry through the physical damage caused by gall formation, leading to rots. They have not been shown to cause direct damage to the enlarged storage roots, but plants can have reduced height if there was loss of enlarged root weight.<ref>{{cite journal |author1=Caveness, F. E. |date=1982 |title=Root-knot nematodes as parasites of cassava |journal=IITA Research Briefs|volume=3 |issue=2 |pages=2–3}}</ref>
▲
; Insects
===Harvesting===▼
[[File:A Cassava Pest of the Southeastern Nigeria.jpg|thumb|upright|Grasshoppers, here on cassava in Nigeria, are secondary pests of cassava.<ref name="Alvarez 2012"/>]]
▲
===Postharvest handling and storage===▼
▲=== Harvesting ===
Cassava deteriorates after harvest, when the the tubers are first cut. The healing mechanism produces [[coumaric acid]], which oxidizes and blackens the tubers, making them inedible after a few days. This deterioration is related to the accumulation of [[reactive oxygen species]] initiated by cyanide release during mechanical harvesting. Cassava shelf life may be increased up to three weeks by overexpressing a cyanide-insensitive alternative oxidase, which suppressed ROS by 10-fold.<ref>{{cite journal |last1=Zidenga |first1=T |display-authors=etal |year=2012 |title=Extending cassava root shelf life via reduction of reactive oxygen species production |journal=[[Plant Physiology (journal)|Plant Physiology]]|volume=159 |issue=4 |pages=1396–1407 |doi=10.1104/pp.112.200345 |pmc=3425186 |pmid=22711743}}</ref> Post-harvest deterioration is a major obstacle to the export of cassava. Fresh cassava can be preserved like potato, using [[thiabendazole]] or bleach as a fungicide, then wrapping in plastic, freezing, or applying a wax coating.<ref>{{cite web |title=Storage and processing of roots and tubers in the tropics |url=http://www.fao.org/docrep/x5415e/x5415e04.htm |access-date=4 May 2016 |publisher=[[U.N. Food and Agriculture Organization]] |archive-date=22 April 2016 |archive-url=https://web.archive.org/web/20160422120308/http://www.fao.org/docrep/x5415e/x5415e04.htm |url-status=live }}</ref>▼
Before harvest, the leafy stems are removed. The harvest is gathered by pulling up the base of the stem and cutting off the roots.<ref name=CIAT/>
▲Cassava deteriorates after harvest, when
While alternative methods for controlling post-harvest deterioration have been proposed, such as preventing reactive oxygen species effects by using plastic bags during storage and transport, coating the roots with wax, or freezing roots, such strategies have proved to be economically or technically impractical, leading to [[plant breeding|breeding]] of cassava varieties with improved durability after harvest, achieved by different mechanisms.<ref name="gmr">{{cite journal |last1=Venturini |first1=M. T |last2=Santos |first2=L. R |last3=Vildoso |first3=C. I |last4=Santos |first4=V. S |last5=Oliveira |first5=E. J |year=2016 |title=Variation in cassava germplasm for tolerance to post-harvest physiological deterioration |journal=[[Genetics and Molecular Research]] |volume=15 |issue=2 |doi=10.4238/gmr.15027818 |pmid=27173317 |doi-access=free}}</ref><ref name="gmr" /><ref name="cs">{{cite journal |last1=Morante |first1=N. |last2=Sánchez |first2=T. |last3=Ceballos |first3=H. |last4=Calle |first4=F. |last5=Pérez |first5=J. C. |last6=Egesi |first6=C. |last7=Cuambe |first7=C. E. |last8=Escobar |first8=A. F. |last9=Ortiz |first9=D. |last10=Chávez |first10=A. L. |last11=Fregene |first11=M. |year=2010 |title=Tolerance to Postharvest Physiological Deterioration in Cassava Roots |journal=[[Crop Science (journal)|Crop Science]]|volume=50 |issue=4 |pages=1333–1338 |doi=10.2135/cropsci2009.11.0666}}</ref> One approach used [[gamma ray]]s to try to silence a gene involved in triggering deterioration; another strategy selected for plentiful [[carotenoid]]s, [[antioxidant]]s which may help to reduce oxidization after harvest.<ref name="cs" />
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== Production ==
{{see also|Tapioca#Production}}▼
{|class="wikitable" style="float:right; clear:right; width:14em; text-align:center; margin-right:1em;"
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|colspan=2|<small>Source: [[FAOSTAT]] of the [[United Nations]]<ref name="faostat">{{cite web|url=http://faostat3.fao.org/browse/Q/QC/E|title=Cassava production in 2022, Crops/World Regions/Production Quantity/Year from pick lists|date=2024|publisher=UN [[Food and Agriculture Organization Corporate Statistical Database]] (FAOSTAT)|access-date=29 May 2024}}</ref></small>
|}
In 2022, world production of cassava root was 330 million tonnes, led by [[Nigeria]] with 18% of the total (table). Other major growers were [[Democratic Republic of the Congo]] and [[Thailand]].
Cassava is the third-largest source of carbohydrates in food in the tropics, after [[rice]] and [[maize]].<ref name="FAO">{{cite web|title=Cassava|publisher=[[Food and Agriculture Organization]] of the United Nations (FAO) |url=http://www.fao.org/ag/agp/agpc/gcds/|access-date=24 November 2011|archive-date=18 November 2016|archive-url=https://web.archive.org/web/20161118061344/http://www.fao.org/ag/agp/agpc/gcds/}}</ref><ref name="Claude-Denis-1990">{{cite journal |author1=Fauquet Claude |author2=Fargette Denis |year=1990 |title=African Cassava Mosaic Virus: Etiology, Epidemiology, and Control |url=http://www.apsnet.org/publications/PlantDisease/BackIssues/Documents/1990Articles/PlantDisease74n06_404.pdf |journal=[[Plant Disease (journal)|Plant Disease]] |publisher=[[American Phytopathological Society]] (APS) |volume=74 |issue=6 |pages=404–411 |doi=10.1094/pd-74-0404 |access-date=10 January 2011 |archive-date=9 August 2017 |archive-url=https://web.archive.org/web/20170809004240/http://www.apsnet.org/publications/PlantDisease/BackIssues/Documents/1990Articles/PlantDisease74n06_404.pdf |url-status=live }}</ref><ref name="Afedraru-2019">{{cite web |last=Afedraru |first=Lominda |title=Uganda to launch innovative gene-edited cassava research |website=Alliance for Science |date=2019-01-31 |url=http://allianceforscience.cornell.edu/blog/2019/01/uganda-launch-innovative-gene-edited-cassava-research/ |access-date=2021-08-15 |archive-date=15 August 2021 |archive-url=https://web.archive.org/web/20210815171521/https://allianceforscience.cornell.edu/blog/2019/01/uganda-launch-innovative-gene-edited-cassava-research/ |url-status=live }}</ref> making it is an important [[staple food|staple]]; more than 500 million people depend on it.<ref>{{cite web|title=Dimensions of Need: An atlas of food and agriculture|publisher=United Nations [[Food and Agriculture Organization]] (FAO) |year=1995|url=http://www.fao.org/docrep/u8480e/U8480E01.htm|access-date=23 November 2011|archive-date=24 November 2016|archive-url=https://web.archive.org/web/20161124095410/http://www.fao.org/docrep/u8480e/U8480E01.htm|url-status=live}}</ref> It offers the advantage of being exceptionally [[drought-tolerant]], and able to grow productively on poor soil. Cassava grows well within 30° of the equator, where it can be produced at up to {{convert|2000|m|ft|sigfig=1|abbr=on}} above sea level, and with {{convert|50|to|5000|mm|in|sigfig=1|abbr=on}} of rain per year. These environmental tolerances suit it to conditions across much of South America and Africa.<ref>{{cite journal |last=Cock |first=James H. |title=Cassava |journal=The Crop Productivity Symposium, IRRI, los Banos, Philippines |date=September 1980 |pages=1–33 |url=https://cgspace.cgiar.org/server/api/core/bitstreams/0bb17dbb-ef96-4ea9-9ca6-100ae47f0503/content}} reprinted as a chapter in ''Crop physiology case histories for major crops''. [[Academic Press]], 2021, pages 588-633.</ref>
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Cassava yields a large amount of food energy per unit area of land per day – {{cvt|250,000|kcal/ha|kJ/ha|order=flip}}, as compared with {{cvt|156,000|kcal/ha|kJ/ha|order=flip}} for rice, {{cvt|110,000|kcal/ha|kJ/ha|order=flip}} for wheat and {{cvt|200,000|kcal/ha|kJ/ha|order=flip}} for maize.<ref>{{Cite journal |last=El-Sharkawy |first=Mabrouk A. |date=1 August 1993 |title=Drought-tolerant Cassava for Africa, Asia, and Latin America |url=http://academic.oup.com/bioscience/article/43/7/441/249072/Droughttolerant-Cassava-for-Africa-Asia-and-Latin |journal=[[BioScience]] |language=en |volume=43 |issue=7 |pages=441–451 |doi=10.2307/1311903 |jstor=1311903 |issn=1525-3244 |access-date=19 April 2020 |archive-date=21 January 2022 |archive-url=https://web.archive.org/web/20220121012151/https://academic.oup.com/bioscience/article/43/7/441/249072/Droughttolerant-Cassava-for-Africa-Asia-and-Latin |url-status=live}}</ref>
Cassava, [[Yam (vegetable)|yams]] (''[[Dioscorea]]'' spp.), and [[sweet potato]]es (''Ipomoea batatas'') are important sources of food in the tropics. The cassava plant gives the third-highest yield of [[carbohydrates]] per cultivated area among crop plants, after [[sugarcane]] and [[sugar beet]]s.<ref>{{cite web |url=http://www.gardeningplaces.com/articles/nutrition-per-hectare1.htm |title=Nutrition per Hectare for Staple Crops |website=GardeningPlaces.com |archive-url=https://web.archive.org/web/20160609084139/http://www.gardeningplaces.com/articles/nutrition-per-hectare1.htm |archive-date=9 June 2016 |url-status=live}}</ref> Cassava plays a particularly important role in agriculture in developing countries, especially in [[sub-Saharan Africa]], because it does well on poor soils and with low rainfall, and because it is a perennial that can be harvested as required. Its wide harvesting window allows it to act as a famine reserve and is invaluable in managing labor schedules. It offers flexibility to resource-poor farmers because it serves as either a subsistence or a cash crop.<ref>{{Cite journal |last1=Stone |first1=G. D. |year=2002 |title=Both Sides Now |journal=[[Current Anthropology]]|volume=43 |issue=4 |pages=611–630 |doi=10.1086/341532 |s2cid=18867515}}</ref> Worldwide, 800 million people depend on cassava as their primary food staple.<ref>{{cite book |url=http://www.fao.org/3/a-i3278e.pdf |title=Save and Grow: Cassava |date=2013 |publisher=[[Food and Agriculture Organization]] |isbn=978-92-5-107641-5 |location=[[Rome]] |page=iii |access-date=27 October 2016 |archive-date=23 November 2016 |archive-url=https://web.archive.org/web/20161123042441/http://www.fao.org/3/a-i3278e.pdf |url-status=live }}</ref>▼
==Toxicity==▼
▲== Toxicity ==
Cassava roots, peels and leaves are dangerous to eat raw because they contain two [[Cyanogenic glycoside|cyanogenic glucosides]], [[linamarin]] and [[lotaustralin]]. These are decomposed by [[linamarase]], a naturally occurring [[enzyme]] in cassava, liberating [[hydrogen cyanide]].<ref name="cereda">{{Cite journal |last1=Cereda |first1=M. P. |last2=Mattos |first2=M. C. Y. |year=1996 |title=Linamarin: the Toxic Compound of Cassava |journal=Journal of Venomous Animals and Toxins|volume=2 |pages=6–12 |doi=10.1590/S0104-79301996000100002 |doi-access=free}}</ref> Cassava varieties are often categorized as either bitter or sweet, signifying the presence or absence of toxic levels of cyanogenic glucosides, respectively. The so-called sweet (more accurately non-bitter) cultivars can produce as little as 20 milligrams of [[cyanide]] per kilogram of fresh roots, whereas bitter ones may produce more than 50 times as much (1 g/kg). Cassavas grown during [[drought]] are especially high in these toxins.<ref>{{cite journal |author1=Aregheore E. M. |author2=Agunbiade O. O. |year=1991 |title=The toxic effects of cassava (''Manihot esculenta'' Crantz) diets on humans: a review|journal=Veterinary and Human Toxicology|volume=33 |issue=3 |pages=274–275 |pmid=1650055}}</ref><ref>{{cite journal |author1=White W. L. B. |author2=Arias-Garzon D. I. |author3=McMahon J. M. |author4=Sayre R. T. |year=1998 |title=Cyanogenesis in Cassava, The Role of Hydroxynitrile Lyase in Root Cyanide Production |journal=[[Plant Physiol.]] |volume=116 |issue=4 |pages=1219–1225 |doi=10.1104/pp.116.4.1219 |pmc=35028 |pmid=9536038}}</ref> A dose of 25 mg of pure cassava cyanogenic glucoside, which contains 2.5 mg of cyanide, is sufficient to kill a rat.<ref>{{cite journal |year=2004 |title=Opinion of the Scientific Panel on Food Additives, Flavourings, Processing Aids and Materials in Contact with Food (AFC) on hydrocyanic acid in flavourings and other food ingredients with flavouring properties |url=http://www.efsa.europa.eu/en/efsajournal/pub/105 |journal=[[EFSA Journal]] |volume=105 |pages=1–28 |access-date=6 April 2013 |archive-date=29 September 2015 |archive-url=https://web.archive.org/web/20150929055532/http://www.efsa.europa.eu/en/efsajournal/pub/105 |url-status=live }}</ref> Excess cyanide residue from improper preparation causes goiters and acute cyanide poisoning, and is linked to ataxia (a neurological disorder affecting the ability to walk, also known as ''[[konzo]]'').<ref name="fao.org" /> It has also been linked to tropical fibrocalcific [[pancreatitis]] in humans, leading to chronic pancreatitis.<ref>{{cite journal |author=Bhatia E |year=2002 |title=Tropical calcific pancreatitis: strong association with SPINK1 trypsin inhibitor mutations |journal=[[Gastroenterology (journal)|Gastroenterology]]|volume=123 |issue=4 |pages=1020–1025 |doi=10.1053/gast.2002.36028 |pmid=12360463|doi-access=free }}</ref><ref name="bbc-2019">{{cite web |last1=Harford |first1=Tim |date=September 4, 2019 |title=How do people learn to cook a poisonous plant safely? |url=https://www.bbc.com/news/business-48859333 |access-date=4 September 2019 |work=BBC News |archive-date=4 September 2019 |archive-url=https://web.archive.org/web/20190904023704/https://www.bbc.com/news/business-48859333 |url-status=live }}</ref>▼
▲Cassava roots, peels and leaves are dangerous to eat raw because they contain
Symptoms of acute cyanide intoxication appear four or more hours after ingesting raw or poorly processed cassava: vertigo, vomiting, [[goiter]], [[ataxia]], partial paralysis, collapse, and death.<ref name="promedmail-4799579">{{cite web|url=https://www.promedmail.org/post/4799579|title=CASSAVA POISONING – VENEZUELA|date=29 January 2017|access-date=29 January 2017|publisher=[[ProMED-mail]]|archive-date=2 February 2017|archive-url=https://web.archive.org/web/20170202061550/https://www.promedmail.org/post/4799579}}</ref><ref>{{cite web |title=Cassava poisoning was integral to Episode 177 of Series 17 of the BBC drama 'Doctors' |publisher=BBC |date=5 February 2016 |url=https://www.bbc.co.uk/iplayer/episode/b06zn3j8/doctors-series-17-177-a-taste-of-home |access-date=13 February 2018 |archive-date=8 February 2016 |archive-url=https://web.archive.org/web/20160208105645/http://www.bbc.co.uk/iplayer/episode/b06zn3j8/doctors-series-17-177-a-taste-of-home |url-status=live }}</ref><ref>{{Cite journal |last1=Soto-Blanco |first1=Benito |last2=Górniak |first2=Silvana Lima |date=2010-07-01 |title=Toxic effects of prolonged administration of leaves of cassava (''Manihot esculenta'' Crantz) to goats |journal=Experimental and Toxicologic Pathology|language=en |volume=62 |issue=4 |pages=361–366 |doi=10.1016/j.etp.2009.05.011 |pmid=19559583 |bibcode=2010EToxP..62..361S |issn=0940-2993}}</ref><ref>{{Cite journal |last1=Suharti |first1=Sri |last2=Oktafiani |first2=Hafni |last3=Sudarman |first3=Asep |last4=Baik |first4=Myunggi |last5=Wiryawan |first5=Komang Gede |date=2021-12-01 |title=Effect of cyanide-degrading bacteria inoculation on performance, rumen fermentation characteristics of sheep fed bitter cassava (''Manihot esculenta'' Crantz) leaf meal|journal=Annals of Agricultural Sciences|language=en |volume=66 |issue=2 |pages=131–136 |doi=10.1016/j.aoas.2021.09.001 |s2cid=244191058 |issn=0570-1783|doi-access=free }}</ref> It can be treated easily with an injection of [[thiosulfate]] (which makes sulfur available for the patient's body to detoxify by converting the poisonous cyanide into thiocyanate).<ref name="fao.org"/>
Like many other roots and tubers, both bitter and sweet varieties of cassava contain [[antinutrient|antinutritional]] factors and toxins, with the bitter varieties containing much larger amounts.<ref name="fao.org">{{cite book |publisher=[[Food and Agriculture Organization]] of the United Nations (FAO) |year=1990 |title=Roots, tubers, plantains and bananas in human nutrition |chapter=Ch. 7 Toxic substances and antinutritional factors |chapter-url=http://www.fao.org/docrep/t0207e/T0207E08.htm#Cassava%20toxicity |place=Rome |isbn=9789251028629|url-access=registration |url=https://archive.org/details/rootstubersplant0000unse}}</ref> The more toxic varieties of cassava have been used in some places as [[famine food]] during times of [[food insecurity]].<ref name="promedmail-4799579" /><ref name="fao.org"/> For example, during the [[shortages in Venezuela]] in the late 2010s, dozens of deaths were reported due to Venezuelans resorting to eating bitter cassava in order to curb starvation.<ref>{{cite news |last1=Castro |first1=Maolis |date=6 March 2017 |title=La yuca amarga alimenta la muerte en Venezuela |language=es |work=[[El País]] |url=https://elpais.com/internacional/2017/03/05/america/1488744764_611719.html |access-date=25 February 2018 |archive-date=12 February 2018 |archive-url=https://web.archive.org/web/20180212184804/https://elpais.com/internacional/2017/03/05/america/1488744764_611719.html |url-status=live }}</ref><ref>{{cite news |date=22 February 2018 |title=Estragos de la crisis: Ocho niños han muerto en Aragua por consumir yuca amarga |language=es-ES |work=[[La Patilla]] |url=https://www.lapatilla.com/site/2018/02/22/estragos-de-la-crisis-ocho-ninos-han-muerto-en-aragua-por-consumir-yuca-amarga/ |access-date=25 February 2018 |archive-date=23 February 2018 |archive-url=https://web.archive.org/web/20180223022727/https://www.lapatilla.com/site/2018/02/22/estragos-de-la-crisis-ocho-ninos-han-muerto-en-aragua-por-consumir-yuca-amarga/ |url-status=live }}</ref> Cases of cassava poisoning were also documented during the famine accompanying the [[Great Leap Forward]] (1958–1962) in China.<ref>{{cite book |author=Zhou Xun |title=The Great Famine in China, 1958-1962: A Documentary History |publisher=Yale University Press |year=2012 |chapter=Ch. 3 Seasons of death}}</ref> Farmers may select bitter cultivars to reduce crop losses.<ref name="leisa">{{cite magazine |author1=Chiwona-Karltun, Linley |author2=Katundu, Chrissie |author3=Ngoma, James |author4=Chipungu, Felistus |author5=Mkumbira, Jonathan |author6=Simukoko, Sidney |author7=Jiggins, Janice |year=2002 |title=Bitter cassava and women: an intriguing response to food security |magazine=LEISA Magazine |volume=18 |issue=4 |url=https://www.researchgate.net/publication/255687456 |access-date=22 September 2018 |archive-date=22 September 2018 |archive-url=https://web.archive.org/web/20180922101746/https://www.researchgate.net/publication/255687456 |url-status=live }}</ref>
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{{main|Cassava-based dishes}}
There are [[Cassava-based dishes
[[Alcoholic beverage]]s made from cassava include [[cauim]] (Brazil),<ref name="rschwan">{{cite journal |title=Yeast diversity in rice-cassava fermentations produced by the indigenous Tapirapé people of Brazil |author=Rosane F. Schwan |author2=Euziclei G. Almeida |author3=Maria Aparecida G. Souza-Dias |author4=Lene Jespersen |date=September 2007 |volume=7 |pages=966–972 |journal=FEMS Yeast Research |doi=10.1111/j.1567-1364.2007.00241.x |pmid=17697080 |issue=6 |doi-access=free }}</ref> [[kasiri]] (Venezuela, Guyana, Suriname),<ref>{{cite news |last=van Vark |first=Manon |title=Tribal cures for modern ailments, Surinam |date=28 August 1999 |work=BBC News |url=http://news.bbc.co.uk/2/hi/americas/431829.stm |quote=Their staple food is cassava, from which they make cassava bread and brew ''kasiri'', 'cassava beer'.}}</ref> [[parakari]] or kari (Venezuela, Guyana, Surinam),<ref>{{Cite journal |last=Henkel |first=Terry W. |date=2005-03-01 |title=Parakari, an indigenous fermented beverage using amylolytic Rhizopus in Guyana |url=https://doi.org/10.1080/15572536.2006.11832833 |journal=Mycologia |volume=97 |issue=1 |pages=1–11 |doi=10.1080/15572536.2006.11832833 |pmid=16389951 |s2cid=218588548 |issn=0027-5514}}</ref> and [[nihamanchi]] (South America),<ref>{{cite book |last=Howell |first=Edward |title=Enzyme Nutrition: The Food Enzyme Concept |publisher=Avery Publishing Group |year=1995 |isbn=978-0895292216 |url=https://books.google.com/books?id=h9EgG8O7GgIC&dq=Howell+Enzyme&pg=PP1 |page=49}}</ref>
▲[[Alcoholic beverage]]s made from cassava include [[cauim]] (Brazil),<ref name="rschwan">{{cite journal |title=Yeast diversity in rice-cassava fermentations produced by the indigenous Tapirapé people of Brazil |author=Rosane F. Schwan |author2=Euziclei G. Almeida |author3=Maria Aparecida G. Souza-Dias |author4=Lene Jespersen |date=September 2007 |volume=7 |pages=966–972 |journal=FEMS Yeast Research |doi=10.1111/j.1567-1364.2007.00241.x |pmid=17697080 |issue=6 |doi-access=free }}</ref> [[kasiri]] (Venezuela, Guyana, Suriname),<ref>{{cite news |last=van Vark |first=Manon |title=Tribal cures for modern ailments, Surinam |date=28 August 1999 |work=BBC News |url=http://news.bbc.co.uk/2/hi/americas/431829.stm |quote=Their staple food is cassava, from which they make cassava bread and brew ''kasiri'', 'cassava beer'.}}</ref> [[parakari]] or kari (Venezuela, Guyana, Surinam),<ref>{{Cite journal |last=Henkel |first=Terry W. |date=2005-03-01 |title=Parakari, an indigenous fermented beverage using amylolytic Rhizopus in Guyana |url=https://doi.org/10.1080/15572536.2006.11832833 |journal=Mycologia |volume=97 |issue=1 |pages=1–11 |doi=10.1080/15572536.2006.11832833 |pmid=16389951 |s2cid=218588548 |issn=0027-5514}}</ref> and [[nihamanchi]] (South America),<ref>{{cite book |last=Howell |first=Edward |title=Enzyme Nutrition: The Food Enzyme Concept |publisher=Avery Publishing Group |year=1995 |isbn=978-0895292216 |url=https://books.google.com/books?id=h9EgG8O7GgIC&dq=Howell+Enzyme&pg=PP1 |page=49}}</ref>
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=== Preparation of bitter cassava ===
A safe processing method known as the "wetting method" is to mix the cassava flour with water into a thick paste, spread it in a thin layer over a basket and then let it stand for five hours at 30 °C in the shade.<ref name="fca">{{cite journal |last1=Bradbury |first1=J.H. |date=2006 |title=Simple wetting method to reduce cyanogen content of cassava flour |url=http://biology-assets.anu.edu.au/hosted_sites/CCDN/papers/Howard_06.pdf |journal=[[Journal of Food Composition and Analysis]] |volume=19 |issue=4 |pages=388–393 |doi=10.1016/j.jfca.2005.04.012 |access-date=23 March 2018 |archive-date=5 February 2015 |archive-url=https://web.archive.org/web/20150205063243/http://biology-assets.anu.edu.au/hosted_sites/CCDN/papers/Howard_06.pdf |url-status=live }}</ref> In that time, about 83% of the cyanogenic [[glycoside]]s are broken down by the [[linamarase]]; the resulting hydrogen cyanide escapes to the atmosphere, making the flour safe for consumption the same evening.<ref name="fca" />
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A traditional method used by the [[Lucayan people|Lucayans]] to detoxify
In Guyana the traditional [[cassareep]] is made from
=== Nutrition ===▼
▲===Nutrition===
{{Nutritional value
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}}
Raw cassava is 60% water, 38% [[carbohydrates]], 1% [[protein]], and has negligible [[fat]] (table).<ref name="fao">{{cite web |title=The Global Cassava Development Strategy|year=2004 |author=Tewe, Olumide O.|publisher=[[U.N. Food and Agriculture Organization]] |url=http://www.fao.org/docrep/007/j1255e/j1255e00.htm|access-date=24 November 2011|archive-date=19 January 2012|archive-url=https://web.archive.org/web/20120119045952/http://www.fao.org/docrep/007/j1255e/j1255e00.htm|url-status=live}}</ref> In a {{convert|100|g|oz|frac=2|abbr=off|adj=on}} reference serving, raw cassava provides {{convert|160|kcal|kJ|order=flip|abbr=off}} of [[food energy]] and 23% of the [[Daily Value]] (DV) of [[vitamin C]], but otherwise has no [[micronutrients]] in significant content (i.e. above 10% of the relevant DV)
=== Biofuel ===
===Animal feed===
Cassava tubers and hay are used worldwide as animal feed.
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=== Laundry starch ===
{{further|Laundry starch}}
Cassava is used in laundry products, especially as starch to stiffen shirts and other garments.<ref>{{Cite web |title=Tapioca or Cassava |url=https://www.botanischetuinen.nl/en/plant_en/5818/tapioca-or-cassava |access-date=2022-03-30 |website=www.botanischetuinen.nl |archive-date=20 April 2023 |archive-url=https://web.archive.org/web/20230420125913/https://www.botanischetuinen.nl/en/plant_en/5818/tapioca-or-cassava |url-status=live }}</ref>▼
▲Cassava is used in laundry products, especially as [[laundry starch|starch]] to stiffen shirts and other garments.<ref>{{Cite web |title=Tapioca or Cassava |url=https://www.botanischetuinen.nl/en/plant_en/5818/tapioca-or-cassava |access-date=2022-03-30 |website=www.botanischetuinen.nl |archive-date=20 April 2023 |archive-url=https://web.archive.org/web/20230420125913/https://www.botanischetuinen.nl/en/plant_en/5818/tapioca-or-cassava |url-status=live }}</ref>
▲{{see also|Tapioca#Production}}
▲Cassava, [[Yam (vegetable)|yams]] (''[[Dioscorea]]'' spp.), and [[sweet potato]]es (''Ipomoea batatas'') are important sources of food in the tropics. The cassava plant gives the third-highest yield of [[carbohydrates]] per cultivated area among crop plants, after [[sugarcane]] and [[sugar beet]]s.<ref>{{cite web |url=http://www.gardeningplaces.com/articles/nutrition-per-hectare1.htm |title=Nutrition per Hectare for Staple Crops |website=GardeningPlaces.com |archive-url=https://web.archive.org/web/20160609084139/http://www.gardeningplaces.com/articles/nutrition-per-hectare1.htm |archive-date=9 June 2016 |url-status=live}}</ref> Cassava plays a particularly important role in agriculture in developing countries, especially in [[sub-Saharan Africa]], because it does well on poor soils and with low rainfall, and because it is a perennial that can be harvested as required. Its wide harvesting window allows it to act as a famine reserve and is invaluable in managing labor schedules. It offers flexibility to resource-poor farmers because it serves as either a subsistence or a cash crop.<ref>{{Cite journal |last1=Stone |first1=G. D. |year=2002 |title=Both Sides Now |journal=[[Current Anthropology]]|volume=43 |issue=4 |pages=611–630 |doi=10.1086/341532 |s2cid=18867515}}</ref> Worldwide, 800 million people depend on cassava as their primary food staple.<ref>{{cite book |url=http://www.fao.org/3/a-i3278e.pdf |title=Save and Grow: Cassava |date=2013 |publisher=[[Food and Agriculture Organization]] |isbn=978-92-5-107641-5 |location=[[Rome]] |page=iii |access-date=27 October 2016 |archive-date=23 November 2016 |archive-url=https://web.archive.org/web/20161123042441/http://www.fao.org/3/a-i3278e.pdf |url-status=live }}</ref>
==See also==
|