Augur buzzard: Difference between revisions

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{{Short description|Species of bird}}
{{speciesbox
| image = Augur buzzard (Buteo augur).jpg
| image_caption = [[Ethiopia]]
| image_upright = 1.2
| status = LC
| status_system = IUCN3.1
| status_ref = <ref name="iucn IUCNstatus 11 November 2021">{{IUCNcite iucn |author=BirdLife International |iddate=227320192016 |title=''Buteo augur'' |assessorvolume=BirdLife International2016 |assessor-linkpage=BirdLife Internationale.T22732019A95040751 |versiondoi=201310.22305/IUCN.UK.2016-3.RLTS.T22732019A95040751.en |year=2012 |accessdateaccess-date=2611 November 20132021}}</ref>
| genus = Buteo
| species = augur
| authority = ([[Eduard Rüppell|Rüppell]], 1836)
| range_map = 1427 - = Augur Buzzard ebird data map.png
| range_map_caption = Global map of [[eBird]] reports{{leftlegend|#007F00|Year-Round Range|outline=gray}}{{leftlegend|#E0CF01|Summer Range|outline=gray}}{{leftlegend|#0080FF|Winter Range|outline=gray}}
}}
 
The '''augur buzzard''' ('''''Buteo augur''''') is a fairly large [[Africa]]n [[bird of prey]]. This species Theis taxonomydistinct onin thistypical speciesadult isplumage confusing,for withits someblackish taxonomistsback, consideringwhitish thisunderside speciesand orange-red tail, thewhile juvenile augur buzzards are generally rather brown in colour; however a [[jackalPolymorphism (biology)|dark buzzardmorph]] is known, andwhich causes the [[Archerbird's buzzard]]entire body to bebecome thedarker. sameThis member of the ''[[superspeciesButeo]].'' genus Manyis taxonomistsdistributed considerin themseveral allparts toof bethe distinct,central havingand differentsouthern callsAfrica, differentnormally homebeing rangesfound andfrom variations[[Ethiopia]] into plumagesouthern [[Angola]] and central [[Namibia]]. It is resident and [[bird migration|non-migratory]] throughout its range. This is a species of mountains (most typically at about {{convert|2000&nbsp;|m|ft|abbr=on}} altitude, but up to {{convert|5000&nbsp;|m|ft|abbr=on}}), and adjacent savannah and grassland. ItThis is residenta andtypical buteonine raptor, being a [[birdGeneralist migrationand specialist species|non-migratorygeneralist predator]] throughoutwhich itstends range.to prefer It is normally found fromsmall [[Ethiopiamammal]]s tosupplemented southernby [[Angolareptile]]s and central [[Namibiabird]]s among various prey items.<ref Thisname= speciesFerguson-Lees>Ferguson-Lees, isJ., also& Christie, D. A. (2001). ''Raptors of the [[Seattleworld''. Seahawks]]Houghton liveMifflin mascotHarcourt.</ref>
 
==Taxonomy==
The taxonomy on this species is not settled, with some taxonomists considering this species, the [[jackal buzzard]], and [[Archer's buzzard]] to be within the same [[superspecies]]. As noted by taxonomists, each species is fairly distinct, having different calls and variations in plumage. While the augur and jackal have rarely been considered actually [[Biological specificity|conspecific]], the Archer's buzzard is sometimes considered improbably as a subspecies of the jackal buzzard despite a number of outward distinctions and having a rather [[Allopatric speciation|allopatric]] and restricted distribution. The three species may be classified as belonging to a [[species complex]].<ref>Kruckenhauser, L., Haring, E., Pinsker, W., Riesing, M. J., Winkler, H., Wink, M., & Gamauf, A. (2004). ''Genetic vs. morphological differentiation of Old World buzzards (genus Buteo, Accipitridae)''. Zoologica Scripta, 33(3), 197-211.</ref><ref>James, A. H. (1986). ''Review of taxonomic characters in African buzzards (genus Buteo)''. Beaufortia, 36(1), 1-12.</ref><ref>Brooke, R. K. (1975). ''The taxonomic relationship of Buteo rufofuscus and B. augur''. Bulletin of the British Ornithologists’ Club, 95, 152-154.</ref><ref>Kemp, A. C. and G. M. Kirwan (2020). ''Archer's Buzzard (Buteo archeri)'', version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA.</ref> In 2022, the International Ornithological Congress tentatively assigned Archer's buzzard as being a [[subspecies]] of the augur buzzard (''Buteo augur archeri''). However, it may simply represent a [[Polymorphism (biology)|color morph]].<ref name=":0">{{Cite web |title=Hoatzin, New World vultures, Secretarybird, raptors – IOC World Bird List |url=https://www.worldbirdnames.org/new/bow/raptors/ |access-date=2022-08-24 |language=en-US}}</ref>
 
Presently, two subspecies are recognized:<ref name=":0" />
 
* ''B. a. augur'' <small>([[Eduard Rüppell|Rüppell]], 1836)</small> - Ethiopia south to Zimbabwe, Angola south to central Namibia
* ''B. a. archeri'' <small>[[William Lutley Sclater|Sclater, WL]], 1918</small> ([[Archer's buzzard]]) - montane northern [[Somalia]]
 
==Description==
[[File:Augur buzzard kisoroAugurbussard-Serengeti.jpg|thumb|left|A melanisticAn augur buzzard in flight[[Serengeti National Park]].]]
Augur buzzards are one of two larger ''[[Buteo]]'' species endemicnative to Africa, alongside their cousins, the similarly sized [[jackal buzzard]]. Adults measure about {{convert|48|to|60|cm|in|abbr=on}} with a large wingspan of {{convert|120|to|149150|cm|ftin|0|abbr=on}}.<ref name= Ferguson-Lees>Ferguson-Lees, J., & Christie, D. A. (2001). ''Raptors of the world''. Houghton Mifflin Harcourt.</ref> Males weigh from {{convert|880|to|1160|g|lb|abbr=on}} while females weigh from {{convert|1100|to|1330|g|lb|abbr=on}}.<ref>''CRC Handbook of Avian Body Masses, 2nd Edition'' by John B. Dunning Jr. (Editor). CRC Press (2008), {{ISBN|978-1-4200-6444-5}}.</ref> A small sample of 5 augur buzzard weighed an average of {{convert|973.2|g|lb|abbr=on}} while 22 birds averaged {{convertcvt|131.5|cm|ftin|0|abbr=on}} in wingspan.<ref>Mendelsohn, J. M., Kemp, A. C., Biggs, H. C., Biggs, R., & Brown, C. J. (1989). ''Wing areas, wing loadings and wing spans of 66 species of African raptors''. Ostrich, 60(1), 35-42.</ref> The adult augur buzzard is strikingly plumaged and essentially unmistakable if seen well. It is an almost black brown above with a [[rufous]] tail that stands out strongly in contrast. The primary flight feathers are blackish and the secondaries off-white, both barred with black. Below the chin and around the throat is mainly white, and the rest of the underparts and the underwing coverts are rich rufous. The flight feathers from below are white, tipped with black to form a dark trailing edge to the wing.
 
The juvenile augur buzzard is mainly brown above and rufous brown below and on the tail. It can be confused with wintering [[Common buzzard|steppe buzzard]], but the augur is considerably larger and bulkier with broader wings and a heavier flight style and an unbarred undertail. Although not as dark as the adult on the back and upperwing coverts, it is usually noticeably darker than a juvenile steppe buzzard. The adult augur buzzard has white underparts and underwings. The female has black on the lower throat. Juveniles are brown above and buff below, the underparts later becoming white. Juveniles are similar to juvenile [[jackal buzzard]]s but are generally much paler below with bolder carpal patches and more clearly barred secondaries and tail. There is one additional subspecies of augur buzzard, endemic to northern [[Somalia]], ''B. a. archeri''. It is mildly smaller than other augur buzzards with extensive rufous mottling about the black-brown above and is almost entirely rufous below but for a small amount of rufous and black about the throat. There is a melanistic form of augur buzzard, all black, except for grey and white flight feathers that are barred black and contrast strongly with the black center and a chestnut tail. About 10% of birds are melanistic, but the proportion rises in forested areas with high rainfall to as much as 50% in some areas. A somewhat similar melanistic morph of [[jackal buzzard]] is also known and these birds can very hard to distinguish, perhaps only told apart by the stronger barring on the melanistic augur pale flight feathers. Dark morph [[long-legged buzzard]]s may also be confused for melanistic augur buzzards but are clearly more slender in the wing, less blackish on the body and lack the bold rufous tail. There are no melanistic individuals known in the ''B. a. archeri'' subspecies.<ref name= Ferguson-Lees/><ref name= Lendrum>Lendrum, A. L. (1979). ''The augur buzzard in the Matopos, Rhodesia''. Ostrich, 50(4), 203-214.</ref>
 
==Range and habitat==
[[File:Augur Buzzard 2.jpg|thumb|right|Gatamayu Forest - Kenya]]
The augur buzzard is found in eastern and southwestern Africa. Despite its erratic looking-seeming distribution, it is often common in its range. The augur buzzard is found from eastern [[Sudan]] and [[Ethiopia]]<ref name="Meheretu2019"/> (also northern [[Somalia]], inhabited by the subspecies ''B. a. archeri'') down through the northeast of the [[Democratic Republic of the Congo]], [[Uganda]], [[Kenya]], parts of [[Tanzania]] into [[Zambia]], [[Malawi]], [[Zimbabwe]], [[Mozambique]], western [[Angola]] and west-central [[Namibia]].<ref name="iucn status 11 November 2021" IUCN/><ref name= Ferguson-Lees/> Despite its wide range in southern Africa it appears to only occur in [[South Africa]] as a vagrant.<ref>Layard, E. L. (1884). ''The birds of South Africa''. Bernard Quartich.</ref> The augur buzzard is found in [[Woodland edge|open]] or light [[Woodland|wooded]] [[Upland and lowland|upland]] areas but can also range into lowland deserts[[desert]]s at sea- level (such as in [[Namibia]]) and some more [[Mountain ecosystems|mountainous, precipitous areas]] of eastern Africa. By preference, theyThey seem to prefer to hunt in [[Plateau|elevated]] [[Savanna|savanna grasslands or]], high [[moorland]], [[Arable land|cropland]]<ref name="Meheretu2019">{{cite book |last1=Meheretu Yonas |last2=Leirs |first2=H |title=''Raptor perch sites for biological control of agricultural pest rodents''. In: Nyssen J., Jacob, M., Frankl, A. (Eds.). Geo-trekking in Ethiopia's Tropical Mountains - The Dogu'a Tembien District |date=2019 |publisher=SpringerNature |isbn=978-3-030-04954-6 |url=https://www.springer.com/gp/book/9783030049546}}</ref> sometimes into [[Dry tropical forest|open forest]] or deserts[[Semi-arid climate|desert edges]] as well. Augur buzzards in [[east Africa]] usually live between {{convert|400|and|4600|m|ft|abbr=on}} elevation but normally occur above {{convert|1500|m|ft|abbr=on}} and have been recorded living at {{convert|5000|m|ft|abbr=on}} in [[Ethiopia]].<ref name= Ferguson-Lees/>
 
==Behaviour==
Pairs have noisy aerial displays, including outside the breeding season. Their call is quite different from their cousin, the jackal buzzard, and most other birds of prey, being a harsh, resonant [[crow]]-like ''a-kow a-kow a-kow'' or ''a-ung a-ung a-ung'', drawn out as aerial display escalates into a longer, higher-pitched ''a-waaa a-waaa a-waaa''.<ref name= Brown>Brown, Leslie and Amadon, Dean (1986) ''Eagles, Hawks and Falcons of the World''. The Wellfleet Press. {{ISBN|978-1555214722}}.</ref> Pairs of augur buzzards usually mate for life but some [[Animal sexual behaviour#Polygamy|polygamy]] has been reported in the species.<ref>Paviour, J. (2013). ''Key factors that influence breeding performance in raptors''. The Plymouth Student Scientist, 6(1), 398-411.</ref> The large (up to {{convertcvt|1|m|ft|abbrdisp=onsqbr}} wide) stick nest is built in a tree or on a crag, and is often reused and enlarged in subsequent seasons. On average two (sometimes only one and rarely three) creamy or bluish white eggs are laid and incubated by the female only, although food is brought to her on the nest by the male. The eggs hatch in about 40 days, and after a further 56–6056&ndash;60 days the chicks can attempt flight. At 70 days they become independent of the nest, but young birds may then be seen with the adult pair for some time. As is the case in other tropical raptors, the clutch size is relatively smaller and the reproductive cycle is relatively longer than in related species found in the temperate zones.<ref name= "Ferguson-Lees" />
[[File:Augurbussard-Serengeti.jpg|thumb|left|An augur buzzard in [[Serengeti National Park]].]]
Pairs have noisy aerial displays, including outside the breeding season. Their call is quite different from their cousin, the jackal buzzard, and most other birds of prey, being a harsh, resonant [[crow]]-like ''a-kow a-kow a-kow'' or ''a-ung a-ung a-ung'', drawn out as aerial display escalates into a longer, higher-pitched ''a-waaa a-waaa a-waaa''.<ref name= Brown>Brown, Leslie and Amadon, Dean (1986) ''Eagles, Hawks and Falcons of the World''. The Wellfleet Press. {{ISBN|978-1555214722}}.</ref> Pairs of augur buzzards usually mate for life but some [[polygamy]] has been reported in the species.<ref>Paviour, J. (2013). ''Key factors that influence breeding performance in raptors''. The Plymouth Student Scientist, 6(1), 398-411.</ref> The large (up to {{convert|1|m|ft|abbr=on}} wide) stick nest is built in a tree or on a crag, and is often reused and enlarged in subsequent seasons. On average two (sometimes only one and rarely three) creamy or bluish white eggs are laid and incubated by the female only, although food is brought to her on the nest by the male. The eggs hatch in about 40 days, and after a further 56–60 days the chicks can attempt flight. At 70 days they become independent of the nest, but young birds may then be seen with the adult pair for some time. As is the case in other tropical raptors, the clutch size is relatively smaller and the reproductive cycle is relatively longer than in related species found in the temperate zones.<ref name= Ferguson-Lees/>
 
The diet of the augur buzzard is quite varied and opportunistic, as is typical of most ''[[Buteo]]'' species. It catches most of its prey on the ground, either by still-hunting from perch or swooping down from a soaring flight or, occasionally, from a hovering flight. They may also forage on the ground for both [[insect]]s and small [[vertebrate]]s. The primary foods for augur buzzards seem to include either small, terrestrial [[mammal]]s or [[reptile]]s, chiefly [[snake]]s and [[lizard]]s. Other prey may include small ground [[bird]]s (and sometimes the nestlings, fledglings or unwary adults of varied birds), [[insect]]s, and road-kill. In [[Zimbabwe]], 59% of the diet was comprised of reptiles while the remainder was mostly mammalian, leadled by [[Otomys|vlei rats]]. At one nest site there, lizards made up 35% of the foods and snakes 46%.<ref>Irwin, M. P. S. (1981). ''The birds of Zimbabwe''. Quest Pub.</ref> In [[Tanzania]], the stomach contents of augur buzzards were similarly comprisedconsisted mostly of assorted [[rat]] species and [[lizard]]s.<ref name="Loveridge">Loveridge, A. (1923, December). ''Notes on East African Birds (chiefly nesting habits and endo‐parasitesendo-parasites) collected 1920–1923''. In Proceedings of the Zoological Society of London (Vol. 93, No. 4, pp. 899-921). Oxford, UK: Blackwell Publishing Ltd.</ref> In the above [[Zimbabwe]] study, the most often taken reptiles recorded the [[giant plated lizard]]s and [[common flat lizard]]s but could extend to larger and more dangerous prey such as [[Nile monitor]]s (though doubtfully large adults) and highly venomous snakes such as [[Bitis arietans|puff adder]]s, [[Causus|night adders]] and [[Mozambique spitting cobra]]s.<ref name= Steyn>Steyn, P. (1983). ''Birds of prey of southern Africa: Their identification and life histories''. Croom Helm, Beckenham (UK). 1983.</ref><ref name= Loveridge/> Elsewhere [[Spalacidae|mole-rat]]s may be preferred, such as in [[Kenya]], and these are likely hunted largely from flight as their tendency to stay in the cover of tall grasses makes them difficult to still-hunt.<ref>Virani, M. Z. (1999). ''The breeding ecology and behaviour of the augur buzzard Buteo augur in relation to different land-uses in the southern Lake Naivasha area, Kenya'' (Doctoral dissertation, Biology).</ref> Occasionally larger prey are hunted including [[francolin]]s, [[Chicken|domestic chickens]], [[hare]]s and [[hyrax]]es, although other than rare cases mainly the juveniles of these prey types are targeted (especially in the case of hyraxes[[Cape hyrax]]es).<ref name= Ferguson-Lees/><ref name= Lendrum/><ref>Young, T. P., & Matthew, R. E. (1993). ''Alpine vertebrates of Mount Kenya, with particular notes on the rock hyrax''. East Africa Natural History Society.</ref><ref>Barry, R. E., & Mundy, P. J. (2015). ''Fluctuations in bush and rock hyrax (Hyracoidea: Procaviidae) abundances over a 13-year period in the Matopos, Zimbabwe''. African Journal of Wildlife Research, 45(1), 17-27.</ref><ref>Hockey P. A. R., Dean, W. R. J. & Ryan, P. G. (2005). ''Roberts - Birds of southern Africa, VIIth ed''. The Trustees of the John Voelcker Bird Book Fund, Cape Town.</ref>
 
==Popular culture==
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==Gallery==
<gallery widths="200px200" heights="150px150">
File:009 Augur buzzard in flight in the Ngorongoro Crater Photo by Giles Laurent.jpg|Augur buzzard in flight with top feathers view, [[Ngorongoro Conservation Area]]
Image:Augur Buzzard, Lake Manyara.jpg|[[Lake Manyara]], [[Tanzania]]
ImageFile:Augur buzzardBuzzard, Lake kisoroManyara.jpg|in flight near [[KisoroLake Manyara]], [[UgandaTanzania]]
File:Augur buzzard kisoro.jpg|in flight near [[Kisoro]], [[Uganda]]
Image:Augur Buzzard (Buteo augur) RWD.jpg|Tanzania
ImageFile:Augur Buzzard RWD1(Buteo augur) RWD.jpg|Carolina Raptor CenterTanzania
File:Augur Buzzard in flightRWD1.jpg|Dark-morphCarolina Raptor Center
ImageFile:Augur Buzzard (Buteoin augur) RWDflight.jpg|TanzaniaDark-morph
</gallery>
 
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==External links==
{{Commons category|Buteo augur}}
{{Wikispecies|Buteo augur}}
* Augur buzzard - [http://sabap2.adu.org.za/docs/sabap1/153.pdf Species text in The Atlas of Southern African Birds]
 
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{{Taxonbar|from=Q760623}}
 
{{DEFAULTSORT:buzzard, augur}}
[[Category:Buteo|augur buzzard]]
[[Category:Birds of prey of Sub-Saharan Africa]]
[[Category:Birds of East Africa]]
[[Category:Birds described in 1836|augur buzzard]]
[[Category:Taxa named by Eduard Rüppell|augur buzzard]]