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'''''Equus lambei''''', [[common name|commonly known as]] the '''Yukon horse''' or '''Yukon wild horse''',<ref name="Forsten 1992"/><ref name="Forsten 1986"/> is an [[Extinction|extinct]] [[species]] of the [[genus]] ''[[Equus (genus)|Equus]]''. ''Equus lambei'' ranged across [[North America]] until approximately 10,000 years ago. Based on recent examinations of the mtDNA of ''Equus lambei'' remains, scientists have concluded that ''E. lambei'' was probably much like the extinct [[tarpan]], also known as the Eurasian wild horse, and the living [[Przewalski's horse]].<sup>[6][8]</sup> A partial carcass of ''Equus lambei'' is on display at the [[Yukon Beringia Interpretive Centre]] in [[Whitehorse, Yukon]].<ref name="Yukon Beringia Interpretive Centre"/>
'''''Equus lambei''''', [[common name|commonly known as]] the '''Yukon horse''' or '''Yukon wild horse''',<ref name="Forsten 1992"/><ref name="Forsten 1986"/> is an [[Extinction|extinct]] [[species]] of the [[genus]] ''[[Equus (genus)|Equus]]''. ''Equus lambei'' ranged across [[North America]] until approximately 10,000 years ago. Based on recent examinations of the mtDNA of ''Equus lambei'' remains, scientists have concluded that ''E. lambei'' was probably much like the extinct [[tarpan]], also known as the Eurasian wild horse, and the living [[Przewalski's horse]].<ref name="Burke and Cinq-Mars 1998" /><ref name="MacFadden 1992" /> A partial carcass of ''Equus lambei'' is on display at the [[Yukon Beringia Interpretive Centre]] in [[Whitehorse]], [[Yukon]].<ref name="Yukon Beringia Interpretive Centre"/>


==Description==
==Description==
Evidence from ''E. lambei'' mtDNA has shown that ''Equus lambei'' is a close relative of the modern [[wild horse]], including the [[domestic horse]], ''[[Horse|Equus caballus]]''.<ref name="Harington YBIC"/><ref name="CMoN"/><ref name="Forsten 1986"/> Controversy still surrounds ''E.&nbsp;lambei'' and the divergence of other similar extinct horse species. Through examinations of the bones and teeth of ''Equus lambei'', many similarities can be seen with the modern horse ''Equus caballus''. There is also a strong resemblance to ''E.&nbsp;lambei'' in the metapodials of ''E.&nbsp;caballus przewalskii'' and the phalanges of ''E.&nbsp;caballus lenensis''.<ref name="Forsten 1986"/>
Evidence from ''E. lambei'' mtDNA has shown that ''Equus lambei'' is a close relative of the modern [[wild horse]], including the [[horse|domestic horse]], ''[[Horse|Equus caballus]]''.<ref name="Harington YBIC"/><ref name="CMoN"/><ref name="Forsten 1986"/> Controversy still surrounds ''E.&nbsp;lambei'' and the divergence of other similar extinct horse species. Through examinations of the bones and teeth of ''Equus lambei'', many similarities can be seen with the modern horse ''Equus caballus''. There is also a strong resemblance to ''E.&nbsp;lambei'' in the metapodials of ''E.&nbsp;caballus przewalskii'' and the phalanges of ''E.&nbsp;caballus lenensis''.<ref name="Forsten 1986"/>


Metrical and morphological studies of horse teeth from the Bluefish Caves confirm the close similarity between ''Equus lambei'' with wild and domestic horses alive today.<ref name="Burke and Cinq-Mars 1998"/> ''E.&nbsp;lambei'' is a ''caballus'' horse, not an [[Asinus|ass]].<ref name="Forsten 1986"/> Among living horses, the Yukon horse most closely resembles the Przewalski's horse (''Equus caballus przewalskii'') from Mongolia<ref name="Forsten 1986"/> (now believed to be extinct in the wild) particularly in size and proportions. However, the upper foot bones ([[metapodial]]s) of ''Equus lambei'' are slender compared to Przewalski's horse.<ref name="Forsten 1986"/> The bones of ''E.&nbsp;lambei'' also closely resemble the proximal phalanges of ''E.&nbsp;caballus lenensis'', an extinct subspecies from the late Pleistocene of Siberia.<ref name="MacFadden 1992"/>
Metrical and morphological studies of horse teeth from the Bluefish Caves confirm the close similarity between ''Equus lambei'' with wild and domestic horses alive today.<ref name="Burke and Cinq-Mars 1998"/> ''E.&nbsp;lambei'' is a ''caballus'' horse, not an [[Asinus|ass]].<ref name="Forsten 1986"/> Among living horses, the Yukon horse most closely resembles the Przewalski's horse (''Equus caballus przewalskii'') from Mongolia<ref name="Forsten 1986"/> (once extinct in the wild<ref>{{Cite journal |date=2014-10-09 |title=Equus ferus: King, S.R.B., Boyd, L., Zimmermann, W. & Kendall, B.E. |doi=10.2305/iucn.uk.2015-2.rlts.t41763a45172856.en |doi-access=free }}</ref>) particularly in size and proportions. However, the upper foot bones ([[metapodial]]s) of ''Equus lambei'' are slender compared to Przewalski's horse.<ref name="Forsten 1986"/> The bones of ''E.&nbsp;lambei'' also closely resemble the proximal phalanges of ''E.&nbsp;caballus lenensis'', an extinct subspecies from the late Pleistocene of Siberia.<ref name="MacFadden 1992"/>

== Taxonomy ==
The taxonomy of Pleistocene North American horses has historically been confused, with many named species that probably represent synonyms of other named species. ''Equus lambei'' is one of the better supported species due to being based on diagnostic type material. Some authors have suggested that ''Equus lambei'' may be synonymous with the larger species ''[[Equus scotti]],'' which may be in turn synonymous with ''Equus ferus''.''<ref name=":0">{{Cite journal |last1=Cirilli |first1=Omar |last2=Machado |first2=Helena |last3=Arroyo-Cabrales |first3=Joaquin |last4=Barrón-Ortiz |first4=Christina I. |last5=Davis |first5=Edward |last6=Jass |first6=Christopher N. |last7=Jukar |first7=Advait M. |last8=Landry |first8=Zoe |last9=Marín-Leyva |first9=Alejandro H. |last10=Pandolfi |first10=Luca |last11=Pushkina |first11=Diana |last12=Rook |first12=Lorenzo |last13=Saarinen |first13=Juha |last14=Scott |first14=Eric |last15=Semprebon |first15=Gina |date=2022-08-24 |title=Evolution of the Family Equidae, Subfamily Equinae, in North, Central and South America, Eurasia and Africa during the Plio-Pleistocene |journal=Biology |language=en |volume=11 |issue=9 |pages=1258 |doi=10.3390/biology11091258 |doi-access=free |issn=2079-7737 |pmc=9495906 |pmid=36138737}}</ref>'' However, North American and Eurasian horses split around 800,000 years ago following the dispersal of the latter out of North America, and have been largely genetically separate, with some interbreeding following the split.<ref>{{Cite journal |last1=Vershinina |first1=Alisa O. |last2=Heintzman |first2=Peter D. |last3=Froese |first3=Duane G. |last4=Zazula |first4=Grant |last5=Cassatt-Johnstone |first5=Molly |last6=Dalén |first6=Love |last7=Der Sarkissian |first7=Clio |last8=Dunn |first8=Shelby G. |last9=Ermini |first9=Luca |last10=Gamba |first10=Cristina |last11=Groves |first11=Pamela |last12=Kapp |first12=Joshua D. |last13=Mann |first13=Daniel H. |last14=Seguin-Orlando |first14=Andaine |last15=Southon |first15=John |date=December 2021 |title=Ancient horse genomes reveal the timing and extent of dispersals across the Bering Land Bridge |url=https://onlinelibrary.wiley.com/doi/10.1111/mec.15977 |journal=Molecular Ecology |language=en |volume=30 |issue=23 |pages=6144–6161 |doi=10.1111/mec.15977 |pmid=33971056 |bibcode=2021MolEc..30.6144V |hdl=10037/24463 |s2cid=234360028 |issn=0962-1083 |hdl-access=free |access-date=2024-01-18 |archive-date=2024-01-11 |archive-url=https://web.archive.org/web/20240111180947/https://onlinelibrary.wiley.com/doi/10.1111/mec.15977 |url-status=live }}</ref>


==Natural history==
==Natural history==
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Based on the fossil records discovered in the Yukon, ''Equus lambei'' is believed to have been a small, slender, caballoid horse (about {{convert|4|ft|abbr=on}} tall), with a broad skull and a relatively long protocones.<ref name="Burke and Cinq-Mars 1998"/><ref name="MacFadden 1992"/>
Based on the fossil records discovered in the Yukon, ''Equus lambei'' is believed to have been a small, slender, caballoid horse (about {{convert|4|ft|abbr=on}} tall), with a broad skull and a relatively long protocones.<ref name="Burke and Cinq-Mars 1998"/><ref name="MacFadden 1992"/>

In August 2018, a 40,000 year-old [[foal]] was discovered in the [[Batagaika crater]], [[Sakha Republic|Yakutia]] in a well preserved condition.<ref name="Sky News 2008"/>


==Social structure==
==Social structure==
Line 38: Line 39:
== Habitat and diet ==
== Habitat and diet ==
Both family and bachelor herds were non-selective grazers that forged mostly in a savannah-like regions.<ref name="Berger 1986"/> These horses fed mostly on grasses, sedges, poppies, mustards, and other flowers such as buttercups and roses. ''Equus lambei's'' preferred environment is believed to have been a woodland with sparse clumps of trees. Overall, ''E.&nbsp;lambei'' is considered to have been resistant to varying climatic conditions, although most individuals of this species seemed to have died in the winter season. It is also likely that the ''Equus lambei'' was susceptible to wolf predation.<ref name="Burke and Cinq-Mars 1998"/><ref name="Waring 1983"/>
Both family and bachelor herds were non-selective grazers that forged mostly in a savannah-like regions.<ref name="Berger 1986"/> These horses fed mostly on grasses, sedges, poppies, mustards, and other flowers such as buttercups and roses. ''Equus lambei's'' preferred environment is believed to have been a woodland with sparse clumps of trees. Overall, ''E.&nbsp;lambei'' is considered to have been resistant to varying climatic conditions, although most individuals of this species seemed to have died in the winter season. It is also likely that the ''Equus lambei'' was susceptible to wolf predation.<ref name="Burke and Cinq-Mars 1998"/><ref name="Waring 1983"/>

==See also==
*[[Yakutian horse]]
*[[Evolution of the horse]]


== References ==
== References ==
{{reflist|refs=
{{reflist|30em|refs=
<ref name="Forsten 1992">{{cite journal|last=Forstén|first=Ann|title=Mitochondrial-DNA time-table and the evolution of ''Equus'': Comparison of molecular and paleontological evidence|journal=Annales Zoologici Fennici|volume=28|number=3/4|pages=301–309|year=1992|jstor=23735454}}</ref>
<ref name="Forsten 1992">{{cite journal|last=Forstén|first=Ann|title=Mitochondrial-DNA time-table and the evolution of ''Equus'': Comparison of molecular and paleontological evidence|journal=Annales Zoologici Fennici|volume=28|number=3/4|pages=301–309|year=1992|jstor=23735454}}</ref>
<ref name="Forsten 1986">{{cite journal|last=Forsten|first=Ann|title=''Equus lambei'' Hay, the Yukon Wild Horse, Not Ass.|journal=Journal of Mammalogy|volume=67|number=2|date=May 15, 1986|pages=422–23|doi=10.2307/1380904|jstor=1380904}}</ref>
<ref name="Forsten 1986">{{cite journal|last=Forsten|first=Ann|title=''Equus lambei'' Hay, the Yukon Wild Horse, Not Ass.|journal=Journal of Mammalogy|volume=67|number=2|date=May 15, 1986|pages=422–23|doi=10.2307/1380904|jstor=1380904}}</ref>
<ref name="Berger 1986">Berger, J. 1986. Wild horses of the Great Basin. Chicago, Illinois: University of Chicago Press.</ref>
<ref name="Berger 1986">Berger, J. 1986. Wild horses of the Great Basin. Chicago, Illinois: University of Chicago Press.</ref>
<ref name="Burke and Cinq-Mars 1998">{{cite journal|last=Burke|first=A.|last2=Cinq-Mars|first2=J.|title=Paleoethological Reconstruction and Taphonomy of ''Equus lambei'' from the Bluefish Caves, Yukon Territory, Canada|doi=10.14430/arctic1052|journal=Arctic|volume=51|number=2|institution=University of Calgary|date=January 1, 1998|pages=105–115}}</ref>
<ref name="Burke and Cinq-Mars 1998">{{cite journal|last1=Burke|first1=A.|last2=Cinq-Mars|first2=J.|title=Paleoethological Reconstruction and Taphonomy of ''Equus lambei'' from the Bluefish Caves, Yukon Territory, Canada|doi=10.14430/arctic1052|journal=Arctic|volume=51|number=2|institution=University of Calgary|date=January 1, 1998|pages=105–115|url=http://doc.rero.ch/record/15588/files/PAL_E3115.pdf|access-date=January 21, 2023|archive-date=April 3, 2023|archive-url=https://web.archive.org/web/20230403110542/https://doc.rero.ch/record/15588/files/PAL_E3115.pdf|url-status=live}}</ref>
<ref name="MacFadden 1992">MacFadden, B.J. 1992. Fossil horses: Systematics, paleobiology and evolution of the family Equidae. Cambridge: Cambridge University Press.</ref>
<ref name="MacFadden 1992">MacFadden, B.J. 1992. Fossil horses: Systematics, paleobiology and evolution of the family Equidae. Cambridge: Cambridge University Press.</ref>
<ref name="Sky News 2008">{{cite web|url=https://news.sky.com/video/square-video-foal-001mp4-11480400|title=40,000-year-old foal discovered in permafrost|publisher=Sky News|date=August 23, 2018}}</ref>
<ref name="Waring 1983">Waring, G.H. 1983. Horse behavior: The behavioral traits and adaptations of domestic and wild horses, including ponies. Park Ridge, New Jersey: Noyes Publications.</ref>
<ref name="Waring 1983">Waring, G.H. 1983. Horse behavior: The behavioral traits and adaptations of domestic and wild horses, including ponies. Park Ridge, New Jersey: Noyes Publications.</ref>
<ref name="Yukon Beringia Interpretive Centre">{{cite web|url=http://www.beringia.com/exhibit/ice-age-animals/yukon-horse|title=Yukon Horse {{!}} Yukon Beringia Interpretive Centre}}</ref>
<ref name="Yukon Beringia Interpretive Centre">{{cite web|url=http://www.beringia.com/exhibit/ice-age-animals/yukon-horse|title=Yukon Horse {{!}} Yukon Beringia Interpretive Centre|access-date=2020-02-27|archive-date=2020-02-27|archive-url=https://web.archive.org/web/20200227083101/https://www.beringia.com/exhibit/ice-age-animals/yukon-horse|url-status=live}}</ref>
<ref name="Harington YBIC">{{cite web|title = Yukon Horse |author = C.R. Harington |date = August 2002 |publisher = Yukon Beringia Interpretive Centre |url = http://beringia.com/research/horse.html |access-date = 2009-01-21 |archive-url = https://web.archive.org/web/20090227093605/http://www.beringia.com/research/horse.html |archive-date = 2009-02-27}}</ref>
<ref name="Harington YBIC">{{cite web|title = Yukon Horse |author = C.R. Harington |date = August 2002 |publisher = Yukon Beringia Interpretive Centre |url = http://beringia.com/research/horse.html |access-date = 2009-01-21 |archive-url = https://web.archive.org/web/20090227093605/http://www.beringia.com/research/horse.html |archive-date = 2009-02-27}}</ref>
<ref name="CMoN">{{cite web|title = Yukon Horse|publisher = Natural History Notebooks. Canadian Museum of Nature|date = 2009-01-20|url = http://www.nature.ca/notebooks/english/yukass.htm|access-date = 2009-01-21}}</ref>
<ref name="CMoN">{{cite web|title = Yukon Horse|publisher = Natural History Notebooks. Canadian Museum of Nature|date = 2009-01-20|url = http://www.nature.ca/notebooks/english/yukass.htm|access-date = 2009-01-21|archive-date = 2009-02-08|archive-url = https://web.archive.org/web/20090208003011/http://nature.ca/notebooks/english/yukass.htm|url-status = live}}</ref>
}}
}}

==External links==
*[https://www.beringia.com/exhibit/ice-age-animals/yukon-horse Yukon Horse]article at Yukon Beringia Interpretive Center in [[Whitehorse, Yukon]]
{{Equus}}
{{Equus}}
{{Taxonbar|from=Q742335}}
{{Taxonbar|from=Q742335}}


[[Category:Equus (genus)]]
[[Category:Equus (genus)|lambei]]
[[Category:Pleistocene horses]]
[[Category:Pleistocene horses]]
[[Category:Holocene extinctions]]
[[Category:Holocene extinctions]]

Latest revision as of 19:30, 17 June 2024

Yukon horse
Temporal range: Late Pleistocene-Early Holocene
Reconstruction of the Yukon horse, based on a skull
Reconstruction of the Yukon horse, based on a skull
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Equidae
Genus: Equus
Subgenus: incertae sedis
Species:
E. lambei
Binomial name
Equus lambei
Hay, 1917

Equus lambei, commonly known as the Yukon horse or Yukon wild horse,[1][2] is an extinct species of the genus Equus. Equus lambei ranged across North America until approximately 10,000 years ago. Based on recent examinations of the mtDNA of Equus lambei remains, scientists have concluded that E. lambei was probably much like the extinct tarpan, also known as the Eurasian wild horse, and the living Przewalski's horse.[3][4] A partial carcass of Equus lambei is on display at the Yukon Beringia Interpretive Centre in Whitehorse, Yukon.[5]

Description

[edit]

Evidence from E. lambei mtDNA has shown that Equus lambei is a close relative of the modern wild horse, including the domestic horse, Equus caballus.[6][7][2] Controversy still surrounds E. lambei and the divergence of other similar extinct horse species. Through examinations of the bones and teeth of Equus lambei, many similarities can be seen with the modern horse Equus caballus. There is also a strong resemblance to E. lambei in the metapodials of E. caballus przewalskii and the phalanges of E. caballus lenensis.[2]

Metrical and morphological studies of horse teeth from the Bluefish Caves confirm the close similarity between Equus lambei with wild and domestic horses alive today.[3] E. lambei is a caballus horse, not an ass.[2] Among living horses, the Yukon horse most closely resembles the Przewalski's horse (Equus caballus przewalskii) from Mongolia[2] (once extinct in the wild[8]) particularly in size and proportions. However, the upper foot bones (metapodials) of Equus lambei are slender compared to Przewalski's horse.[2] The bones of E. lambei also closely resemble the proximal phalanges of E. caballus lenensis, an extinct subspecies from the late Pleistocene of Siberia.[4]

Taxonomy

[edit]

The taxonomy of Pleistocene North American horses has historically been confused, with many named species that probably represent synonyms of other named species. Equus lambei is one of the better supported species due to being based on diagnostic type material. Some authors have suggested that Equus lambei may be synonymous with the larger species Equus scotti, which may be in turn synonymous with Equus ferus.[9] However, North American and Eurasian horses split around 800,000 years ago following the dispersal of the latter out of North America, and have been largely genetically separate, with some interbreeding following the split.[10]

Natural history

[edit]

Along with steppe bison (Bison priscus), woolly mammoths (Mammuthus primigenius) and caribou (Rangifer tarandus), Equus lambei was one of the most common ice-age species known to occupy the steppe-like grasslands of Eastern Beringia.[6][7] E. lambei can be identified by numerous teeth and bones, and one partial carcass discovered in 1993. This E. lambei carcass yielded a radiocarbon date of 26,280 ± 210 years BP.[6][3] The carcass consisted of a large part of the hide, a few tailbones, one lower leg, and some intestine. The hide retained some long blondish mane and tail hairs, coarse whitish upper body hairs, and dark brown hairs on the lower leg.[6][3][2] Large numbers of E. lambei teeth have been found in archaeological sites in the Yukon.[2][4]

Based on the fossil records discovered in the Yukon, Equus lambei is believed to have been a small, slender, caballoid horse (about 4 ft (1.2 m) tall), with a broad skull and a relatively long protocones.[3][4]

Social structure

[edit]

Specifically, the records indicates that E. lambei had a multi-seasonal presence in the same region as other horse species, and a social structure similar to other wild horses. Family herds included four to ten females with their young and an adult alpha stallion.[3][2] Other males were gathered in less socially stable bachelor herds, and consist from groups of two to four adults.[4][11] These two herds did not often share territories.[12]

Habitat and diet

[edit]

Both family and bachelor herds were non-selective grazers that forged mostly in a savannah-like regions.[12] These horses fed mostly on grasses, sedges, poppies, mustards, and other flowers such as buttercups and roses. Equus lambei's preferred environment is believed to have been a woodland with sparse clumps of trees. Overall, E. lambei is considered to have been resistant to varying climatic conditions, although most individuals of this species seemed to have died in the winter season. It is also likely that the Equus lambei was susceptible to wolf predation.[3][11]

See also

[edit]

References

[edit]
  1. ^ Forstén, Ann (1992). "Mitochondrial-DNA time-table and the evolution of Equus: Comparison of molecular and paleontological evidence". Annales Zoologici Fennici. 28 (3/4): 301–309. JSTOR 23735454.
  2. ^ a b c d e f g h i Forsten, Ann (May 15, 1986). "Equus lambei Hay, the Yukon Wild Horse, Not Ass". Journal of Mammalogy. 67 (2): 422–23. doi:10.2307/1380904. JSTOR 1380904.
  3. ^ a b c d e f g Burke, A.; Cinq-Mars, J. (January 1, 1998). "Paleoethological Reconstruction and Taphonomy of Equus lambei from the Bluefish Caves, Yukon Territory, Canada" (PDF). Arctic. 51 (2). University of Calgary: 105–115. doi:10.14430/arctic1052. Archived (PDF) from the original on April 3, 2023. Retrieved January 21, 2023.
  4. ^ a b c d e MacFadden, B.J. 1992. Fossil horses: Systematics, paleobiology and evolution of the family Equidae. Cambridge: Cambridge University Press.
  5. ^ "Yukon Horse | Yukon Beringia Interpretive Centre". Archived from the original on 2020-02-27. Retrieved 2020-02-27.
  6. ^ a b c d C.R. Harington (August 2002). "Yukon Horse". Yukon Beringia Interpretive Centre. Archived from the original on 2009-02-27. Retrieved 2009-01-21.
  7. ^ a b "Yukon Horse". Natural History Notebooks. Canadian Museum of Nature. 2009-01-20. Archived from the original on 2009-02-08. Retrieved 2009-01-21.
  8. ^ "Equus ferus: King, S.R.B., Boyd, L., Zimmermann, W. & Kendall, B.E." 2014-10-09. doi:10.2305/iucn.uk.2015-2.rlts.t41763a45172856.en. {{cite journal}}: Cite journal requires |journal= (help)
  9. ^ Cirilli, Omar; Machado, Helena; Arroyo-Cabrales, Joaquin; Barrón-Ortiz, Christina I.; Davis, Edward; Jass, Christopher N.; Jukar, Advait M.; Landry, Zoe; Marín-Leyva, Alejandro H.; Pandolfi, Luca; Pushkina, Diana; Rook, Lorenzo; Saarinen, Juha; Scott, Eric; Semprebon, Gina (2022-08-24). "Evolution of the Family Equidae, Subfamily Equinae, in North, Central and South America, Eurasia and Africa during the Plio-Pleistocene". Biology. 11 (9): 1258. doi:10.3390/biology11091258. ISSN 2079-7737. PMC 9495906. PMID 36138737.
  10. ^ Vershinina, Alisa O.; Heintzman, Peter D.; Froese, Duane G.; Zazula, Grant; Cassatt-Johnstone, Molly; Dalén, Love; Der Sarkissian, Clio; Dunn, Shelby G.; Ermini, Luca; Gamba, Cristina; Groves, Pamela; Kapp, Joshua D.; Mann, Daniel H.; Seguin-Orlando, Andaine; Southon, John (December 2021). "Ancient horse genomes reveal the timing and extent of dispersals across the Bering Land Bridge". Molecular Ecology. 30 (23): 6144–6161. Bibcode:2021MolEc..30.6144V. doi:10.1111/mec.15977. hdl:10037/24463. ISSN 0962-1083. PMID 33971056. S2CID 234360028. Archived from the original on 2024-01-11. Retrieved 2024-01-18.
  11. ^ a b Waring, G.H. 1983. Horse behavior: The behavioral traits and adaptations of domestic and wild horses, including ponies. Park Ridge, New Jersey: Noyes Publications.
  12. ^ a b Berger, J. 1986. Wild horses of the Great Basin. Chicago, Illinois: University of Chicago Press.
[edit]