Megalosauroidea: Difference between revisions
No edit summary |
I removed the? Of the Piatnitzkysauridae Tags: Visual edit Mobile edit Mobile web edit |
||
(32 intermediate revisions by 22 users not shown) | |||
Line 2: | Line 2: | ||
{{Automatic taxobox |
{{Automatic taxobox |
||
| name = Megalosauroids |
| name = Megalosauroids |
||
| fossil_range = [[Middle Jurassic]] - [[Late Cretaceous]], {{fossilrange|earliest=179|170|93|latest= |
| fossil_range = [[Middle Jurassic]] - [[Late Cretaceous]], {{fossilrange|earliest=179|170|93|latest=85}}<small>Possible Late [[Toarcian]] and Late [[Santonian]] records</small> |
||
| image = Torvosaurus Museum of Ancient Life 2.jpg |
| image = Torvosaurus Museum of Ancient Life 2.jpg |
||
| image_caption = ''[[Torvosaurus]]'' skeletal mount, [[Museum of Ancient Life]] |
| image_caption = ''[[Torvosaurus]]'' skeletal mount, [[Museum of Ancient Life]] |
||
Line 13: | Line 13: | ||
| subdivision_ranks = Subgroups |
| subdivision_ranks = Subgroups |
||
| subdivision = *?{{extinct}}''[[Sciurumimus]]'' |
| subdivision = *?{{extinct}}''[[Sciurumimus]]'' |
||
*?{{extinct}}''[[Juravenator]]'' |
*?{{extinct}}''[[Juravenator]]'' |
||
*?{{extinct}}''[[ |
*?{{extinct}}''[[Dandakosaurus]]'' |
||
* |
*{{extinct}}''[[Yunyangosaurus]]'' |
||
* {{extinct}}[[Piatnitzkysauridae]] |
|||
* {{extinct}}'''Megalosauria''' |
* {{extinct}}'''Megalosauria''' |
||
** {{extinct}}''[[Streptospondylus]]'' |
** {{extinct}}''[[Streptospondylus]]'' |
||
Line 25: | Line 26: | ||
}} |
}} |
||
'''Megalosauroidea''' (meaning 'great/big lizard forms') is a [[Taxonomic rank|superfamily]] (or [[clade]]) of [[Tetanurae|tetanuran]] theropod [[dinosaur]]s that lived from the [[Middle Jurassic]] to the [[Late Cretaceous]] period. The group is defined as ''[[Megalosaurus|Megalosaurus bucklandii]]'' and all taxa sharing a more recent common ancestor with it than with ''[[Allosaurus fragilis]]'' or ''[[Passer domesticus]]''.<ref name="benson2010" /> Members of the group include ''[[Spinosaurus]]'', ''[[Megalosaurus]]'', and ''[[Torvosaurus]]''. They are possibly paraphyletic in nature. |
'''Megalosauroidea''' (meaning 'great/big lizard forms') is a [[Taxonomic rank|superfamily]] (or [[clade]]) of [[Tetanurae|tetanuran]] theropod [[dinosaur]]s that lived from the [[Middle Jurassic]] to the [[Late Cretaceous]] period. The group is defined as ''[[Megalosaurus|Megalosaurus bucklandii]]'' and all taxa sharing a more recent common ancestor with it than with ''[[Allosaurus fragilis]]'' or ''[[Passer domesticus]]''.<ref name="benson2010" /> Members of the group include ''[[Spinosaurus]]'', ''[[Megalosaurus]]'', and ''[[Torvosaurus]]''. They are possibly paraphyletic in nature with respect to [[Allosauroidea]]. |
||
==Classification== |
==Classification== |
||
The name '''Spinosauroidea''' is sometimes used in place of Megalosauroidea. The superfamily Spinosauroidea was named in 1915 by [[Ernst Stromer]]. It is a synonym of Megalosauroidea in almost all modern phylogenetic analyses, and it is therefore redundant. Spinosauroidea was defined as a [[clade]] in 1998 by [[Paul Sereno]] as the node clade containing the common ancestor of ''Spinosaurus'' and ''Torvosaurus'' and all its descendants. Thomas Holtz in 2004 defined a branch clade with the same name containing all species closer to ''Spinosaurus'' than to ''[[Passer]] domesticus''. The [[International Commission on Zoological Nomenclature|ICZN]] holds that even clade names (which do not yet have any governing body) should be replaced if having a traditional taxon suffix and being synonyms of ranked taxa at or below the superfamily level. The seniority of Megalosauroidea was not followed in most paleontological literature during the 1990s and early 2000s. A series of papers supporting the validity of ''[[Megalosaurus]]'' as a genus, the relationships of megalosauroids, and the placement of "spinosauroids" among them, published between 2008 and 2010 argued that Megalosauroidea was in fact the valid name for the group.<ref name="benson2010">{{cite journal |last=Benson |first=R.B.J. |year=2010 |title=A description of ''Megalosaurus bucklandii'' (Dinosauria: Theropoda) from the Bathonian of the UK and the relationships of Middle Jurassic theropods |journal=Zoological Journal of the Linnean Society |doi= 10.1111/j.1096-3642.2009.00569.x |volume=158 |issue=4 |pages=882–935 |doi-access=free }}</ref> |
The name '''Spinosauroidea''' is sometimes used in place of Megalosauroidea. The superfamily Spinosauroidea was named in 1915 by [[Ernst Stromer]]. It is a synonym of Megalosauroidea in almost all modern phylogenetic analyses, and it is therefore redundant. Spinosauroidea was defined as a [[clade]] in 1998 by [[Paul Sereno]] as the node clade containing the common ancestor of ''Spinosaurus'' and ''Torvosaurus'' and all its descendants. Thomas Holtz in 2004 defined a branch clade with the same name containing all species closer to ''Spinosaurus'' than to ''[[Passer]] domesticus''. The [[International Commission on Zoological Nomenclature|ICZN]] holds that even clade names (which do not yet have any governing body) should be replaced if having a traditional taxon suffix and being synonyms of ranked taxa at or below the superfamily level. The seniority of Megalosauroidea was not followed in most paleontological literature during the 1990s and early 2000s. A series of papers supporting the validity of ''[[Megalosaurus]]'' as a genus, the relationships of megalosauroids, and the placement of "spinosauroids" among them, published between 2008 and 2010 argued that Megalosauroidea was in fact the valid name for the group.<ref name="benson2010">{{cite journal |last=Benson |first=R.B.J. |year=2010 |title=A description of ''Megalosaurus bucklandii'' (Dinosauria: Theropoda) from the Bathonian of the UK and the relationships of Middle Jurassic theropods |journal=Zoological Journal of the Linnean Society |doi= 10.1111/j.1096-3642.2009.00569.x |volume=158 |issue=4 |pages=882–935 |doi-access=free }}</ref> |
||
The classification of megalosauroids follows a study by Benson in 2010. Note that several "wildcard"<ref name="benson2010" /> taxa that are probably megalosauroids were excluded from the final tree, including ''[[Magnosaurus]]'', ''[[Piveteausaurus]]'' and ''[[Streptospondylus]]''.<ref name="benson2010" /> Later, ''[[Magnosaurus]]'' and ''Streptospondylus'' were added in the final tree.<ref name="bensonetal2010">{{Cite journal | author = Benson, R.B.J., Carrano, M.T and Brusatte, S.L. | year = 2010 | title = A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic | journal = Naturwissenschaften | volume = 97 | issue = 1 | pages = 71–78 | doi = 10.1007/s00114-009-0614-x | pmid=19826771 | bibcode=2010NW.....97...71B}} [https://doi.org/10.1007%2Fs00114-009-0614-x Supporting Information]</ref> These are known from remains too fragmentary to be reliably classified.<ref name="benson2010" /> Some of these 'wildcard' taxa, such as ''[[Poekilopleuron]]'' and ''[[Megaraptor]]'', have been considered possible megalosauroids in the past, but the analysis found that they are more likely [[allosauroid]]s.<ref name="bensonetal2010" /> |
The classification of megalosauroids follows a study by Benson in 2010. Note that several "wildcard"<ref name="benson2010" /> taxa that are probably megalosauroids were excluded from the final tree, including ''[[Magnosaurus]]'', ''[[Piveteausaurus]]'' and ''[[Streptospondylus]]''.<ref name="benson2010" /> Later, ''[[Magnosaurus]]'' and ''Streptospondylus'' were added in the final tree.<ref name="bensonetal2010">{{Cite journal | author = Benson, R.B.J., Carrano, M.T and Brusatte, S.L. | year = 2010 | title = A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic | journal = Naturwissenschaften | volume = 97 | issue = 1 | pages = 71–78 | doi = 10.1007/s00114-009-0614-x | pmid=19826771 | bibcode=2010NW.....97...71B| s2cid = 22646156 }} [https://doi.org/10.1007%2Fs00114-009-0614-x Supporting Information]</ref> These are known from remains too fragmentary to be reliably classified.<ref name="benson2010" /> Some of these 'wildcard' taxa, such as ''[[Poekilopleuron]]'' and ''[[Megaraptor]]'', have been considered possible megalosauroids in the past, but the analysis found that they are more likely [[allosauroid]]s.<ref name="bensonetal2010" /> |
||
{{clade| style=font-size:85%;line-height:85% |
{{clade| style=font-size:85%;line-height:85% |
||
Line 45: | Line 46: | ||
|1={{clade |
|1={{clade |
||
|1=''[[Chuandongocoelurus]]'' |
|1=''[[Chuandongocoelurus]]'' |
||
|2=''[[Monolophosaurus]]'' [[File:Monolophosaurus jiangi |
|2=''[[Monolophosaurus]]'' [[File:Monolophosaurus jiangi jmallon (flipped).jpg|100 px]]}} |
||
|2={{clade |
|2={{clade |
||
|label1=[[Spinosauridae]] |
|label1=[[Spinosauridae]] |
||
Line 51: | Line 52: | ||
|1={{clade |
|1={{clade |
||
|1=''[[Baryonyx]]'' |
|1=''[[Baryonyx]]'' |
||
|2=''[[Suchomimus]]'' [[File:Suchomimus |
|2=''[[Suchomimus]]'' <span style="{{MirrorH}}">[[File:Suchomimus tenerensis by PaleoGeek.png|100 px]]</span>}} |
||
|2={{clade |
|2={{clade |
||
|1=''[[Irritator]]'' [[File:Irritator challengeri |
|1=''[[Irritator]]'' <span style="{{MirrorH}}">[[File:Irritator challengeri by PaleoGeek.png|100 px]]</span> |
||
|2=''[[Spinosaurus]]'' [[File:Spinosaurus |
|2=''[[Spinosaurus]]'' <span style="{{MirrorH}}">[[File:Spinosaurus aegyptiacus by PaleoGeek.png|100px]]</span>}} }} |
||
|label2=[[Megalosauridae]] |
|label2=[[Megalosauridae]] |
||
|2={{clade |
|2={{clade |
||
Line 67: | Line 68: | ||
|1=''[[Duriavenator]]'' |
|1=''[[Duriavenator]]'' |
||
|2={{clade |
|2={{clade |
||
|1=''[[Afrovenator]]'' [[File:Afrovenator |
|1=''[[Afrovenator]]'' [[File:Afrovenator Abakensis by PaleoGeek.jpg|100 px]] |
||
|2=''[[Dubreuillosaurus]]'' }} |
|2=''[[Dubreuillosaurus]]'' }} |
||
|3={{clade |
|3={{clade |
||
|1=''[[Megalosaurus]]'' |
|1=''[[Megalosaurus]]'' |
||
|2=''[[Torvosaurus]]'' |
|2=''[[Torvosaurus]]'' [[File:Torvosaurus tanner DBi.jpg|100px]]}} }} }} }} }} }} }} |
||
Carrano, Benson & Sampson (2012) performed much larger [[phylogenetic]] analysis of the [[Tetanurae]] that includes more taxa. They used the clade name '''Megalosauria''' (Bonaparte, 1850) in their analysis and defined it as the node comprising ''Megalosaurus'', ''Spinosaurus'', their most recent common ancestor, and all its descendants. Furthermore, a new megalosauroid family [[Piatnitzkysauridae]] was named to include all megalosauroids more closely related to ''Piatnitzkysaurus'' than to either ''Spinosaurus'' or ''Megalosaurus''. Within Megalosauridae a new subfamily was named, [[Afrovenatorinae]], to include all megalosaurids more closely related to ''Afrovenator'' than to ''Megalosaurus''. Unlike Benson ''et al.'', 2010, they recovered ''[[Poekilopleuron]]'' as an afrovenatorine, while ''[[Xuanhanosaurus]]'' was recovered as the basalmost [[metriacanthosaurid]]. However, the position of these taxa is very unstable, and their exclusion from the analysis gave more resolved and stable cladogram. ''[[Streptospondylus]]'' was also excluded to get more resolved Megalosauridae and Afrovenatorinae. The Chinese tetanuran ''[[Leshansaurus]]'' was included for the first time in a phylogenetic analysis, and was recovered as an afrovenatorine. Both ''[[Chuandongocoelurus]]'' and ''[[Monolophosaurus]]'' were found to be just outside [[Orionides]]. The [[cladogram]] presented here follows that study.<ref name=tetanurans2012>{{Cite journal | last1 = Carrano | first1 = M. T. | last2 = Benson | first2 = R. B. J. | last3 = Sampson | first3 = S. D. | doi = 10.1080/14772019.2011.630927 | title = The phylogeny of Tetanurae (Dinosauria: Theropoda) | journal = Journal of Systematic Palaeontology | volume = 10 | issue = 2 | pages = 211–300| year = 2012 }}</ref> |
Carrano, Benson & Sampson (2012) performed much larger [[phylogenetic]] analysis of the [[Tetanurae]] that includes more taxa. They used the clade name '''Megalosauria''' (Bonaparte, 1850) in their analysis and defined it as the node comprising ''Megalosaurus'', ''Spinosaurus'', their most recent common ancestor, and all its descendants. Furthermore, a new megalosauroid family [[Piatnitzkysauridae]] was named to include all megalosauroids more closely related to ''Piatnitzkysaurus'' than to either ''Spinosaurus'' or ''Megalosaurus''. Within Megalosauridae a new subfamily was named, [[Afrovenatorinae]], to include all megalosaurids more closely related to ''Afrovenator'' than to ''Megalosaurus''. Unlike Benson ''et al.'', 2010, they recovered ''[[Poekilopleuron]]'' as an afrovenatorine, while ''[[Xuanhanosaurus]]'' was recovered as the basalmost [[metriacanthosaurid]]. However, the position of these taxa is very unstable, and their exclusion from the analysis gave more resolved and stable cladogram. ''[[Streptospondylus]]'' was also excluded to get more resolved Megalosauridae and Afrovenatorinae. The Chinese tetanuran ''[[Leshansaurus]]'' was included for the first time in a phylogenetic analysis, and was recovered as an afrovenatorine. Both ''[[Chuandongocoelurus]]'' and ''[[Monolophosaurus]]'' were found to be just outside [[Orionides]]. The [[cladogram]] presented here follows that study.<ref name=tetanurans2012>{{Cite journal | last1 = Carrano | first1 = M. T. | last2 = Benson | first2 = R. B. J. | last3 = Sampson | first3 = S. D. | doi = 10.1080/14772019.2011.630927 | title = The phylogeny of Tetanurae (Dinosauria: Theropoda) | journal = Journal of Systematic Palaeontology | volume = 10 | issue = 2 | pages = 211–300| year = 2012 | s2cid = 85354215 }}</ref> |
||
{{clade| style=font-size:85%;line-height:85% |
{{clade| style=font-size:85%;line-height:85% |
||
|label1=[[Tetanurae]] |
|label1=[[Tetanurae]] |
||
|1={{clade |
|1={{clade |
||
|1=''[[Cryolophosaurus]]'' [[File:Cryolophosaurus |
|1=''[[Cryolophosaurus]]'' [[File:Cryolophosaurus reconstruction (flipped).jpg|100 px]] |
||
|2=''[[Sinosaurus|"Dilophosaurus" sinensis]]'' [[File: |
|2=''[[Sinosaurus|"Dilophosaurus" sinensis]]'' [[File:Dilophosaurus wetherilli (flipped).PNG|100 px]] |
||
|3={{clade |
|3={{clade |
||
|1=''[[Chuandongocoelurus]]'' |
|1=''[[Chuandongocoelurus]]'' |
||
|2=''[[Monolophosaurus]]'' [[File:Monolophosaurus jiangi |
|2=''[[Monolophosaurus]]'' [[File:Monolophosaurus jiangi jmallon (flipped).jpg|100 px]] |
||
|label3=[[Orionides]] |
|label3=[[Orionides]] |
||
|3={{clade |
|3={{clade |
||
|label1=[[Avetheropoda]] |
|label1=[[Avetheropoda]] |
||
|1={{clade |
|1={{clade |
||
|1=[[Allosauroidea]] [[File:Allosaurus |
|1=[[Allosauroidea]] [[File:Allosaurus Revised.jpg|120px]] |
||
|label2=[[Coelurosauria]] |
|label2=[[Coelurosauria]] |
||
|2={{clade |
|2={{clade |
||
|1=''[[Lourinhanosaurus]]'' |
|1=''[[Lourinhanosaurus]]'' |
||
|2=Other coelurosaurians [[File: |
|2=Other coelurosaurians <span style="{{MirrorH}}">[[File:Deinonychus ewilloughby.png|100 px]]</span>}} }} |
||
|label2='''Megalosauroidea''' |
|label2='''Megalosauroidea''' |
||
|2={{clade |
|2={{clade |
||
Line 107: | Line 108: | ||
|1={{clade |
|1={{clade |
||
|1=''[[Baryonyx]]'' |
|1=''[[Baryonyx]]'' |
||
|2=''[[Suchomimus]]'' [[File:Suchomimus |
|2=''[[Suchomimus]]'' <span style="{{MirrorH}}">[[File:Suchomimus tenerensis by PaleoGeek.png|100 px]]</span>}} |
||
|2={{clade |
|2={{clade |
||
|1=''[[Angaturama]]'' |
|1=''[[Angaturama]]'' |
||
|2=''[[Irritator]]'' [[File:Irritator challengeri |
|2=''[[Irritator]]'' <span style="{{MirrorH}}">[[File:Irritator challengeri by PaleoGeek.png|100 px]]</span> |
||
|3=''[[Spinosaurus]]'' [[File:Spinosaurus |
|3=''[[Spinosaurus]]'' <span style="{{MirrorH}}">[[File:Spinosaurus aegyptiacus by PaleoGeek.png|100px]]</span>}} }} |
||
|label3=[[Megalosauridae]] |
|label3=[[Megalosauridae]] |
||
|3={{clade |
|3={{clade |
||
Line 123: | Line 124: | ||
|2={{clade |
|2={{clade |
||
|1=''[[Megalosaurus]]'' |
|1=''[[Megalosaurus]]'' |
||
|2=''[[Torvosaurus]]'' |
|2=''[[Torvosaurus]]'' [[File:Torvosaurus tanner DBi.jpg|100px]]}} }} |
||
|label2=[[Afrovenatorinae]] |
|label2=[[Afrovenatorinae]] |
||
|2={{clade |
|2={{clade |
||
|1=''[[Afrovenator]]'' [[File:Afrovenator |
|1=''[[Afrovenator]]'' [[File:Afrovenator Abakensis by PaleoGeek.jpg|100 px]] |
||
|2={{clade |
|2={{clade |
||
|1=''[[Dubreuillosaurus]]'' |
|1=''[[Dubreuillosaurus]]'' |
||
Line 134: | Line 135: | ||
|2=''[[Piveteausaurus]]'' }} }} }} }} }} }} }} }} }} }} |
|2=''[[Piveteausaurus]]'' }} }} }} }} }} }} }} }} }} }} |
||
In 2019, Rauhut and Pol described ''[[Asfaltovenator|Asfaltovenator vialidadi]]'', a basal allosauroid displaying a mosaic of primitive and derived features seen within [[Tetanurae]]. Their phylogenetic analysis found traditional Megalosauroidea to represent a basal [[evolutionary grade|grade]] of carnosaurs, [[paraphyly|paraphyletic]] with respect to [[Allosauroidea]].<ref name="Rauhut2019">{{Cite journal| |
In 2019, Rauhut and Pol described ''[[Asfaltovenator|Asfaltovenator vialidadi]]'', a basal allosauroid displaying a mosaic of primitive and derived features seen within [[Tetanurae]]. Their phylogenetic analysis found traditional Megalosauroidea to represent a basal [[evolutionary grade|grade]] of carnosaurs, [[paraphyly|paraphyletic]] with respect to [[Allosauroidea]].<ref name="Rauhut2019">{{Cite journal|last1=Rauhut|first1=Oliver W. M.|last2=Pol|first2=Diego|date=2019-12-11|title=Probable basal allosauroid from the early Middle Jurassic Cañadón Asfalto Formation of Argentina highlights phylogenetic uncertainty in tetanuran theropod dinosaurs|journal=Scientific Reports|language=en|volume=9|issue=1|pages=18826|doi=10.1038/s41598-019-53672-7|pmid=31827108|pmc=6906444|bibcode=2019NatSR...918826R |issn=2045-2322}}</ref> |
||
{{clade| style=font-size:100%;line-height:80% |
{{clade| style=font-size:100%;line-height:80% |
||
Line 142: | Line 143: | ||
|1=''[[Monolophosaurus]]''[[File:Monolophosaurus jiangi jmallon (flipped).jpg|100 px]] |
|1=''[[Monolophosaurus]]''[[File:Monolophosaurus jiangi jmallon (flipped).jpg|100 px]] |
||
|2={{clade |
|2={{clade |
||
|1=[[Spinosauridae]][[File:Spinosaurus aegyptiacus.png|100px]]|barbegin1=midnightblue |
|1=[[Spinosauridae]]<span style="{{MirrorH}}">[[File:Spinosaurus aegyptiacus by PaleoGeek.png|100px]]</span>|barbegin1=midnightblue |
||
|2={{clade |
|2={{clade |
||
|1=[[Megalosauridae]][[File:Torvosaurus tanner DBi.jpg| |
|1=[[Megalosauridae]][[File:Torvosaurus tanner DBi.jpg|100px]]|bar1=midnightblue |
||
|label2=[[Allosauroidea]] |
|label2=[[Allosauroidea]] |
||
|2={{clade |
|2={{clade |
||
Line 156: | Line 157: | ||
|2={{clade |
|2={{clade |
||
|1=''[[Allosaurus]]''[[File:Allosaurus Revised.jpg|120px]] |
|1=''[[Allosaurus]]''[[File:Allosaurus Revised.jpg|120px]] |
||
|2=[[Carcharodontosauria]][[File:Carcharodontosaurus.png|120px]]}} }} }} }} }} }} }} }} }} |
|2=[[Carcharodontosauria]]<span style="{{MirrorH}}">[[File:Carcharodontosaurus.png|120px]]</span>}} }} }} }} }} }} }} }} }} |
||
==See also== |
==See also== |
||
Line 168: | Line 169: | ||
{{Taxonbar|from=Q132202}} |
{{Taxonbar|from=Q132202}} |
||
[[Category: |
[[Category:Tetanurans]] |
||
[[Category:Fossil taxa described in 1889]] |
|||
[[Category:Bajocian first appearances]] |
[[Category:Bajocian first appearances]] |
||
[[Category:Cenomanian extinctions]] |
[[Category:Cenomanian extinctions]] |
Revision as of 19:57, 1 July 2024
Megalosauroids Temporal range: Middle Jurassic - Late Cretaceous, Possible Late Toarcian and Late Santonian records
| |
---|---|
![]() | |
Torvosaurus skeletal mount, Museum of Ancient Life | |
![]() | |
Spinosaurus skeletal mount, National Geographic Museum | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Clade: | †Carnosauria (?) |
Superfamily: | †Megalosauroidea Huxley, 1889 |
Type species | |
†Megalosaurus bucklandii Mantell, 1827
| |
Subgroups | |
| |
Synonyms | |
Megalosauroidea (meaning 'great/big lizard forms') is a superfamily (or clade) of tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as Megalosaurus bucklandii and all taxa sharing a more recent common ancestor with it than with Allosaurus fragilis or Passer domesticus.[1] Members of the group include Spinosaurus, Megalosaurus, and Torvosaurus. They are possibly paraphyletic in nature with respect to Allosauroidea.
Classification
The name Spinosauroidea is sometimes used in place of Megalosauroidea. The superfamily Spinosauroidea was named in 1915 by Ernst Stromer. It is a synonym of Megalosauroidea in almost all modern phylogenetic analyses, and it is therefore redundant. Spinosauroidea was defined as a clade in 1998 by Paul Sereno as the node clade containing the common ancestor of Spinosaurus and Torvosaurus and all its descendants. Thomas Holtz in 2004 defined a branch clade with the same name containing all species closer to Spinosaurus than to Passer domesticus. The ICZN holds that even clade names (which do not yet have any governing body) should be replaced if having a traditional taxon suffix and being synonyms of ranked taxa at or below the superfamily level. The seniority of Megalosauroidea was not followed in most paleontological literature during the 1990s and early 2000s. A series of papers supporting the validity of Megalosaurus as a genus, the relationships of megalosauroids, and the placement of "spinosauroids" among them, published between 2008 and 2010 argued that Megalosauroidea was in fact the valid name for the group.[1]
The classification of megalosauroids follows a study by Benson in 2010. Note that several "wildcard"[1] taxa that are probably megalosauroids were excluded from the final tree, including Magnosaurus, Piveteausaurus and Streptospondylus.[1] Later, Magnosaurus and Streptospondylus were added in the final tree.[2] These are known from remains too fragmentary to be reliably classified.[1] Some of these 'wildcard' taxa, such as Poekilopleuron and Megaraptor, have been considered possible megalosauroids in the past, but the analysis found that they are more likely allosauroids.[2]
Megalosauroidea |
| ||||||
Carrano, Benson & Sampson (2012) performed much larger phylogenetic analysis of the Tetanurae that includes more taxa. They used the clade name Megalosauria (Bonaparte, 1850) in their analysis and defined it as the node comprising Megalosaurus, Spinosaurus, their most recent common ancestor, and all its descendants. Furthermore, a new megalosauroid family Piatnitzkysauridae was named to include all megalosauroids more closely related to Piatnitzkysaurus than to either Spinosaurus or Megalosaurus. Within Megalosauridae a new subfamily was named, Afrovenatorinae, to include all megalosaurids more closely related to Afrovenator than to Megalosaurus. Unlike Benson et al., 2010, they recovered Poekilopleuron as an afrovenatorine, while Xuanhanosaurus was recovered as the basalmost metriacanthosaurid. However, the position of these taxa is very unstable, and their exclusion from the analysis gave more resolved and stable cladogram. Streptospondylus was also excluded to get more resolved Megalosauridae and Afrovenatorinae. The Chinese tetanuran Leshansaurus was included for the first time in a phylogenetic analysis, and was recovered as an afrovenatorine. Both Chuandongocoelurus and Monolophosaurus were found to be just outside Orionides. The cladogram presented here follows that study.[3]
Tetanurae |
| ||||||||||||||||||||||||
In 2019, Rauhut and Pol described Asfaltovenator vialidadi, a basal allosauroid displaying a mosaic of primitive and derived features seen within Tetanurae. Their phylogenetic analysis found traditional Megalosauroidea to represent a basal grade of carnosaurs, paraphyletic with respect to Allosauroidea.[4]
Carnosauria | traditional Megalosauroidea | |
See also
References
- ^ a b c d e Benson, R.B.J. (2010). "A description of Megalosaurus bucklandii (Dinosauria: Theropoda) from the Bathonian of the UK and the relationships of Middle Jurassic theropods". Zoological Journal of the Linnean Society. 158 (4): 882–935. doi:10.1111/j.1096-3642.2009.00569.x.
- ^ a b Benson, R.B.J., Carrano, M.T and Brusatte, S.L. (2010). "A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic". Naturwissenschaften. 97 (1): 71–78. Bibcode:2010NW.....97...71B. doi:10.1007/s00114-009-0614-x. PMID 19826771. S2CID 22646156.
{{cite journal}}
: CS1 maint: multiple names: authors list (link) Supporting Information - ^ Carrano, M. T.; Benson, R. B. J.; Sampson, S. D. (2012). "The phylogeny of Tetanurae (Dinosauria: Theropoda)". Journal of Systematic Palaeontology. 10 (2): 211–300. doi:10.1080/14772019.2011.630927. S2CID 85354215.
- ^ Rauhut, Oliver W. M.; Pol, Diego (2019-12-11). "Probable basal allosauroid from the early Middle Jurassic Cañadón Asfalto Formation of Argentina highlights phylogenetic uncertainty in tetanuran theropod dinosaurs". Scientific Reports. 9 (1): 18826. Bibcode:2019NatSR...918826R. doi:10.1038/s41598-019-53672-7. ISSN 2045-2322. PMC 6906444. PMID 31827108.