Abstract
Unbalanced production of proinflammatory cytokines and type I interferons in immune responses may lead to immunopathology; thus, the mechanisms that ensure the beneficial production of proinflammatory cytokines and type I interferons are of particular importance. Here we demonstrate that the phosphatase SHP-1 negatively regulated Toll-like receptor-mediated production of proinflammatory cytokines by inhibiting activation of the transcription factor NF-kappaB and mitogen-activated protein kinase. Simultaneously, SHP-1 increased the production of type I interferon mediated by Toll-like receptors and the helicase RIG-I by directly binding to and inhibiting activation of the kinase IRAK1. Our data demonstrate that SHP-1 contributes to immune homeostasis by balancing the production of proinflammatory cytokines and type I interferons in the innate immune response.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Catalytic Domain / physiology
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Cytokines / biosynthesis
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Homeostasis / immunology
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Immunity, Innate
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Interferon Regulatory Factor-1 / metabolism
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Interferon Type I / biosynthesis*
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Interleukin-1 Receptor-Associated Kinases / antagonists & inhibitors
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Interleukin-1 Receptor-Associated Kinases / chemistry
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Interleukin-1 Receptor-Associated Kinases / metabolism*
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Macrophages, Peritoneal
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Membrane Proteins / immunology
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Membrane Proteins / metabolism*
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Mice
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Mice, Inbred C57BL
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Mitogen-Activated Protein Kinase Kinases / antagonists & inhibitors
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Mitogen-Activated Protein Kinase Kinases / metabolism
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NF-kappa B / antagonists & inhibitors
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NF-kappa B / metabolism
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Nerve Tissue Proteins / immunology
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Nerve Tissue Proteins / metabolism*
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Protein Binding
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Protein Tyrosine Phosphatase, Non-Receptor Type 6 / immunology
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Protein Tyrosine Phosphatase, Non-Receptor Type 6 / metabolism*
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Receptors, Cell Surface
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Signal Transduction
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Toll-Like Receptors / immunology
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Toll-Like Receptors / metabolism*
Substances
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Cytokines
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Interferon Regulatory Factor-1
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Interferon Type I
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Membrane Proteins
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NF-kappa B
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Nerve Tissue Proteins
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Receptors, Cell Surface
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Robo3 protein, mouse
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Toll-Like Receptors
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Interleukin-1 Receptor-Associated Kinases
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Irak1 protein, mouse
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Mitogen-Activated Protein Kinase Kinases
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Protein Tyrosine Phosphatase, Non-Receptor Type 6