Caspase recruitment domain-containing protein 8 (CARD8) negatively regulates NOD2-mediated signaling

J Biol Chem. 2010 Jun 25;285(26):19921-6. doi: 10.1074/jbc.M110.127480. Epub 2010 Apr 12.

Abstract

Caspase activating and recruitment domain 8 (CARD8) has been implicated as a co-regulator of several pro-inflammatory and apoptotic signaling pathways. In the present study, we demonstrate a specific modulation of NOD2-induced signaling by CARD8 in intestinal epithelial cells. We show that CARD8 physically interacts with NOD2 and inhibits nodosome assembly and subsequent signaling upon muramyl-dipeptide stimulation. Furthermore, CARD8 inhibits the direct bactericidal effect of NOD2 against intracellular infection by Listeria monocytogenes. Thus, CARD8 represents a novel molecular switch involved in the endogenous regulation of NOD2-dependent inflammatory processes in epithelial cells.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Acetylmuramyl-Alanyl-Isoglutamine / pharmacology
  • Anti-Bacterial Agents / pharmacology
  • Blotting, Western
  • CARD Signaling Adaptor Proteins / genetics
  • CARD Signaling Adaptor Proteins / metabolism*
  • Caco-2 Cells
  • Cell Line
  • Cell Line, Tumor
  • Colon / enzymology*
  • Crohn Disease / genetics
  • Crohn Disease / metabolism
  • Crohn Disease / pathology
  • Gene Expression Profiling
  • Gentamicins / pharmacology
  • HT29 Cells
  • HeLa Cells
  • Humans
  • Intestinal Mucosa / metabolism
  • Intestinal Mucosa / pathology
  • Intracellular Space / drug effects
  • Intracellular Space / metabolism
  • Intracellular Space / microbiology
  • Listeria monocytogenes / growth & development
  • Luminescent Proteins / genetics
  • Luminescent Proteins / metabolism
  • Male
  • Microscopy, Fluorescence
  • Neoplasm Proteins / genetics
  • Neoplasm Proteins / metabolism*
  • Nod2 Signaling Adaptor Protein / genetics
  • Nod2 Signaling Adaptor Protein / metabolism*
  • Protein Binding
  • RNA Interference
  • Reverse Transcriptase Polymerase Chain Reaction
  • Signal Transduction / drug effects
  • Signal Transduction / genetics
  • Signal Transduction / physiology*
  • Transfection

Substances

  • Anti-Bacterial Agents
  • CARD Signaling Adaptor Proteins
  • CARD8 protein, human
  • Gentamicins
  • Luminescent Proteins
  • NOD2 protein, human
  • Neoplasm Proteins
  • Nod2 Signaling Adaptor Protein
  • Acetylmuramyl-Alanyl-Isoglutamine