Abstract
Targeted inactivation of Runx3 in mouse lung induced mucinous and nonmucinous adenomas and markedly shortened latency of adenocarcinoma formation induced by oncogenic K-Ras. RUNX3 was frequently inactivated in K-RAS mutated human lung adenocarcinomas. A functional genetic screen of a fly mutant library and molecular analysis in cultured cell lines revealed that Runx3 forms a complex with BRD2 in a K-Ras-dependent manner in the early phase of the cell cycle; this complex induces expression of p14(ARF)/p19(Arf) and p21(WAF/CIP). When K-Ras was constitutively activated, the Runx3-BRD2 complex was stably maintained and expression of both p14(ARF) and p21(WAF/CIP) was prolonged. These results provide a missing link between oncogenic K-Ras and the p14(ARF)-p53 pathway, and may explain how cells defend against oncogenic K-Ras.
Copyright © 2013 Elsevier Inc. All rights reserved.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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ADP-Ribosylation Factors / metabolism
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Acetylation
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Adenocarcinoma / metabolism*
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Adenocarcinoma of Lung
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Alveolar Epithelial Cells / physiology
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Animals
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Carcinogenesis / metabolism
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Cell Differentiation
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Cell Line, Tumor
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Core Binding Factor Alpha 3 Subunit / genetics
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Core Binding Factor Alpha 3 Subunit / metabolism*
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Cyclin D1 / metabolism
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Cyclin-Dependent Kinase Inhibitor p21 / metabolism
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Gene Expression
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Gene Knockout Techniques
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HEK293 Cells
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Histone Deacetylases / metabolism
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Humans
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Lung Neoplasms / metabolism*
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Mice
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Mice, Transgenic
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Protein Processing, Post-Translational
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Protein Serine-Threonine Kinases / metabolism
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Proto-Oncogene Proteins / genetics
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Proto-Oncogene Proteins / metabolism
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Proto-Oncogene Proteins p21(ras) / genetics
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Proto-Oncogene Proteins p21(ras) / metabolism
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Repressor Proteins / metabolism
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Respiratory Mucosa / pathology
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Transcription Factors
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ras Proteins / genetics
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ras Proteins / metabolism
Substances
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BRD2 protein, human
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CCND1 protein, human
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CDKN1A protein, human
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Core Binding Factor Alpha 3 Subunit
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Cyclin-Dependent Kinase Inhibitor p21
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KRAS protein, human
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Proto-Oncogene Proteins
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Repressor Proteins
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Runx3 protein, human
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Runx3 protein, mouse
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Transcription Factors
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Cyclin D1
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Protein Serine-Threonine Kinases
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HDAC4 protein, human
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Histone Deacetylases
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ADP-Ribosylation Factors
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Hras protein, mouse
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Proto-Oncogene Proteins p21(ras)
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ras Proteins