Allele-specific induction of IL-1β expression by C/EBPβ and PU.1 contributes to increased tuberculosis susceptibility

PLoS Pathog. 2014 Oct 16;10(10):e1004426. doi: 10.1371/journal.ppat.1004426. eCollection 2014 Oct.

Abstract

Mycobacterium tuberculosis infection is associated with a spectrum of clinical outcomes, from long-term latent infection to different manifestations of progressive disease. Pro-inflammatory pathways, such as those controlled by IL-1β, have the contrasting potential both to prevent disease by restricting bacterial replication, and to promote disease by inflicting tissue damage. Thus, the ultimate contribution of individual inflammatory pathways to the outcome of M. tuberculosis infection remains ambiguous. In this study, we identified a naturally-occurring polymorphism in the human IL1B promoter region, which alters the association of the C/EBPβ and PU.1 transcription factors and controls Mtb-induced IL-1β production. The high-IL-1β expressing genotype was associated with the development of active tuberculosis, the severity of pulmonary disease and poor treatment outcome in TB patients. Higher IL-1β expression did not suppress the activity of IFN-γ-producing T cells, but instead correlated with neutrophil accumulation in the lung. These observations support a specific role for IL-1β and granulocytic inflammation as a driver of TB disease progression in humans, and suggest novel strategies for the prevention and treatment of tuberculosis.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Alleles*
  • CCAAT-Enhancer-Binding Protein-beta / genetics*
  • Cell Line
  • Disease Susceptibility / microbiology*
  • Humans
  • Interferon-gamma / metabolism
  • Interleukin-1beta / genetics*
  • Lung / microbiology
  • Proto-Oncogene Proteins / genetics*
  • Trans-Activators / genetics*
  • Tuberculosis / microbiology*

Substances

  • CCAAT-Enhancer-Binding Protein-beta
  • Interleukin-1beta
  • Proto-Oncogene Proteins
  • Trans-Activators
  • proto-oncogene protein Spi-1
  • Interferon-gamma

Grants and funding

This work was supported by Twelfth-Fifth Mega-Scientific Project on “prevention and treatment of AIDS, viral hepatitis and other infectious diseases” (2012ZX10003002), Natural Science Foundation of China Grant (81273140, 81341128), Natural Science Foundation of Guangdong Grant (S2012040007213), Science and Technology Foundation of Shenzhen Grant (JCYJ20140411111718166) and the Howard Hughes Medical Institute. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.