Activation of PPARγ by 12/15-lipoxygenase during cerebral ischemia-reperfusion injury

Int J Mol Med. 2015 Jan;35(1):195-201. doi: 10.3892/ijmm.2014.1998. Epub 2014 Nov 12.

Abstract

Peroxisome proliferator-activated receptor γ (PPARγ) expression and activity are increased in brain ischemic injury and its agonists have shown potential for brain injury protection. The influence of 12/15-lipoxygenase (12/15-LOX) on the activity of PPARγ in oxygen-glucose deprivation (OGD) and ischemia-reperfusion (I/R) was investigated. A middle cerebral artery occlusion/reperfusion model with Sprague Dawley (SD) rats was established. For I/R intervention, the rats were treated with the 12/15-LOX-derived product 12-hydroxyeicosatetraenoic acid (12-HETE) for 30 min before cerebral artery occlusion. Primary cortical neurons from SD rats were used to establish an OGD cell model. 12-HETE or a 12/15-LOX antisense oligonucleotide (asON-12/15-LOX) was added to OGD-treated neurons. Western blots, immunofluorescence and enzyme-linked immunosorbent assays detected protein. Reverse transcription-polymerase chain reaction analyzed the expression of the PPARγ target genes. PPARγ-DNA binding activity was determined by peroxisome proliferator responsive element luciferase reporter vectors. 12/15-LOX total protein increased significantly with I/R, and expression of 12-HETE was also upregulated. 12-HETE treatment increased PPARγ protein expression and inhibited inducible nitric oxide synthase protein expression, which was upregulated with I/R. PPARγ nuclear protein and 12/15-LOX total protein expression in OGD-treated neurons increased significantly. 12-HETE treatment increased the expression of PPARγ nuclear protein, upregulated the mRNA levels of PPARγ target genes (lipoprotein lipase and acyl-CoA oxidase) and enhanced PPARγ-DNA binding activity. asON-12/15-LOX treatment inhibited 12/15-LOX and PPARγ protein expression and lipoprotein lipase mRNA. Cerebral I/R injury in rats and OGD treatment in neurons promoted 12/15-LOX expression, and 12-HETE activated PPARγ. Therefore, PPARγ can be activated by the 12/15-LOX pathway during cerebral I/R injury.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Arachidonate 12-Lipoxygenase / genetics
  • Arachidonate 12-Lipoxygenase / metabolism*
  • Arachidonate 12-Lipoxygenase / pharmacology
  • Arachidonate 15-Lipoxygenase / genetics
  • Arachidonate 15-Lipoxygenase / metabolism*
  • Arachidonate 15-Lipoxygenase / pharmacology
  • Cerebral Cortex / blood supply
  • Cerebral Cortex / metabolism*
  • Cerebral Cortex / pathology*
  • Disease Models, Animal
  • Enzyme Activation
  • Gene Expression
  • Glucose / metabolism
  • Infarction, Middle Cerebral Artery / complications
  • Male
  • Neurons / drug effects
  • Neurons / metabolism
  • Nitric Oxide Synthase Type II / genetics
  • Nitric Oxide Synthase Type II / metabolism
  • Oxygen / metabolism
  • PPAR gamma / metabolism*
  • Protein Binding
  • Protein Transport
  • Rats
  • Reperfusion Injury / etiology
  • Reperfusion Injury / metabolism*

Substances

  • 12-15-lipoxygenase
  • PPAR gamma
  • Arachidonate 12-Lipoxygenase
  • Arachidonate 15-Lipoxygenase
  • Nitric Oxide Synthase Type II
  • Glucose
  • Oxygen