Memory reconsolidation of an inhibitory avoidance task in mice involves cytosolic ERK2 bidirectional modulation

Neuroscience. 2015 May 21:294:227-37. doi: 10.1016/j.neuroscience.2015.03.019. Epub 2015 Mar 17.

Abstract

Reconsolidation has been defined as the process of memory stabilization after retrieval involving, among others, gene expression regulation and post-translational modifications. Many of these mechanisms are shared with memory consolidation. Here, we studied hippocampal ERK participation on memory reconsolidation of an inhibitory avoidance task in CF-1 mice. We found a retrieval-induced cytosolic ERK2 activation in the hippocampus (HIP) 15 min after memory reactivation, and an inhibition at 45 min. PD098059, a MEK1/2 (MAPK/ERK kinase) inhibitor, administered in the HIP immediately after retrieval impaired memory in a dose-dependent fashion. However, infusions of the highest dose of PD098059 performed 40 min after retrieval enhanced memory in mice trained with a weaker footshock. These results suggest for the first time that ERK2 is involved in memory reconsolidation in a biphasic fashion. Furthermore, the inhibition of ERK could either impair or enhance mice performance depending on ERK state of activation.

Keywords: ERK; MAPK; PD098059; inhibitory avoidance; memory reconsolidation; mice.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Avoidance Learning / physiology*
  • Hippocampus / metabolism
  • MAP Kinase Signaling System / physiology*
  • Male
  • Memory / physiology*
  • Mice
  • Mitogen-Activated Protein Kinase 1 / metabolism*
  • Phosphorylation
  • Protein Processing, Post-Translational / physiology*

Substances

  • Mitogen-Activated Protein Kinase 1