1 UCL Great Ormond Street Institute of Child Health, London, United Kingdom, CoMPLEX: The UCL Centre for Computation, Mathematics and Physics in the Life Sciences and Experimental Biology, University College London, London, United Kingdom UCL Great Ormond Street Institute of Child Health, London, United Kingdom, Great Ormond Street Hospital, London, United Kingdom CoMPLEX: The UCL Centre for Computation, Mathematics and Physics in the Life Sciences and Experimental Biology, University College London, London, United Kingdom, Department of Infectious Disease Epidemiology, Imperial College London, Norfolk Place, United Kingodm.