Dynamics of TERT regulation via alternative splicing in stem cells and cancer cells

PLoS One. 2023 Aug 2;18(8):e0289327. doi: 10.1371/journal.pone.0289327. eCollection 2023.

Abstract

Part of the regulation of telomerase activity includes the alternative splicing (AS) of the catalytic subunit telomerase reverse transcriptase (TERT). Although a therapeutic window for telomerase/TERT inhibition exists between cancer cells and somatic cells, stem cells express TERT and rely on telomerase activity for physiological replacement of cells. Therefore, identifying differences in TERT regulation between stem cells and cancer cells is essential for developing telomerase inhibition-based cancer therapies that reduce damage to stem cells. In this study, we measured TERT splice variant expression and telomerase activity in induced pluripotent stem cells (iPSCs), neural progenitor cells (NPCs), and non-small cell lung cancer cells (NSCLC, Calu-6 cells). We observed that a NOVA1-PTBP1-PTBP2 axis regulates TERT alternative splicing (AS) in iPSCs and their differentiation into NPCs. We also found that splice-switching of TERT, which regulates telomerase activity, is induced by different cell densities in stem cells but not cancer cells. Lastly, we identified cell type-specific splicing factors that regulate TERT AS. Overall, our findings represent an important step forward in understanding the regulation of TERT AS in stem cells and cancer cells.

Publication types

  • Research Support, N.I.H., Extramural

MeSH terms

  • Alternative Splicing
  • Carcinoma, Non-Small-Cell Lung*
  • Heterogeneous-Nuclear Ribonucleoproteins / metabolism
  • Humans
  • Induced Pluripotent Stem Cells* / metabolism
  • Lung Neoplasms*
  • Polypyrimidine Tract-Binding Protein / metabolism
  • Telomerase* / genetics
  • Telomerase* / metabolism

Substances

  • Telomerase
  • TERT protein, human
  • PTBP1 protein, human
  • Heterogeneous-Nuclear Ribonucleoproteins
  • Polypyrimidine Tract-Binding Protein