The p53-mdm-2 autoregulatory feedback loop

Genes Dev. 1993 Jul;7(7A):1126-32. doi: 10.1101/gad.7.7a.1126.

Abstract

The p53 protein can bind to a set of specific DNA sequences, and this may activate the transcription of genes adjacent to these DNA elements. The mdm-2 gene is shown here to contain a p53 DNA-binding site and a genetically responsive element such that expression of the mdm-2 gene can be regulated by the level of wild-type p53 protein. The mdm-2 protein, in turn, can complex with p53 and decrease its ability to act as a positive transcription factor at the mdm-2 gene-responsive element. In this way, the mdm-2 gene is autoregulated. The p53 protein regulates the mdm-2 gene at the level of transcription, and the mdm-2 protein regulates the p53 protein at the level of its activity. This creates a feedback loop that regulates both the activity of the p53 protein and the expression of the mdm-2 gene.

Publication types

  • Research Support, U.S. Gov't, P.H.S.

MeSH terms

  • Animals
  • Base Sequence
  • Binding Sites
  • Cell Line
  • Cell Line, Transformed
  • DNA Mutational Analysis
  • DNA-Binding Proteins / metabolism
  • Feedback
  • Fibroblasts
  • Gene Expression Regulation, Neoplastic*
  • Homeostasis
  • Mice
  • Molecular Sequence Data
  • Mutation
  • Neoplasm Proteins / genetics*
  • Nuclear Proteins*
  • Promoter Regions, Genetic
  • Protein Binding
  • Proto-Oncogene Proteins c-mdm2
  • Proto-Oncogene Proteins*
  • RNA, Messenger / metabolism
  • Rats
  • Temperature
  • Transcription Factors / genetics
  • Transcription, Genetic
  • Transcriptional Activation*
  • Tumor Suppressor Protein p53 / genetics*
  • Tumor Suppressor Protein p53 / physiology

Substances

  • DNA-Binding Proteins
  • Neoplasm Proteins
  • Nuclear Proteins
  • Proto-Oncogene Proteins
  • RNA, Messenger
  • Transcription Factors
  • Tumor Suppressor Protein p53
  • Mdm2 protein, mouse
  • Mdm2 protein, rat
  • Proto-Oncogene Proteins c-mdm2