Abstract
Several members of the tumour-necrosis/nerve-growth factor (TNF/NGF) receptor family activate the transcription factor NF-kappaB through a common adaptor protein, Traf2 (refs 1-5), whereas the interleukin 1 type-I receptor activates NF-kappaB independently of Traf2 (ref. 4). We have now cloned a new protein kinase, NIK, which binds to Traf2 and stimulates NF-kappaB activity. This kinase shares sequence similarity with several MAPKK kinases. Expression in cells of kinase-deficient NIK mutants fails to stimulate NF-kappaB and blocks its induction by TNF, by either of the two TNF receptors or by the receptor CD95 (Fas/Apo-1), and by TRADD, RIP and MORT1/FADD, which are adaptor proteins that bind to these receptors. It also blocked NF-kappaB induction by interleukin-1. Our findings indicate that NIK participates in an NF-kappaB-inducing signalling cascade common to receptors of the TNF/NGF family and to the interleukin-1 type-I receptor.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Adaptor Proteins, Signal Transducing*
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Amino Acid Sequence
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Carrier Proteins / metabolism
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Cell Line
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Cloning, Molecular
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Fas-Associated Death Domain Protein
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Humans
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Interleukin-1 / physiology*
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MAP Kinase Kinase Kinases
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Molecular Sequence Data
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NF-kappa B / metabolism*
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NF-kappaB-Inducing Kinase
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Protein Binding
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Protein Serine-Threonine Kinases / genetics
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Protein Serine-Threonine Kinases / metabolism*
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Proteins / metabolism
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Receptor-Interacting Protein Serine-Threonine Kinases
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Saccharomyces cerevisiae / genetics
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Signal Transduction
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TNF Receptor-Associated Factor 1
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TNF Receptor-Associated Factor 2
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Transfection
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Tumor Necrosis Factor-alpha / physiology*
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fas Receptor / physiology*
Substances
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Adaptor Proteins, Signal Transducing
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Carrier Proteins
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FADD protein, human
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Fas-Associated Death Domain Protein
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Interleukin-1
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NF-kappa B
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Proteins
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TNF Receptor-Associated Factor 1
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TNF Receptor-Associated Factor 2
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Tumor Necrosis Factor-alpha
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fas Receptor
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Protein Serine-Threonine Kinases
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RIPK1 protein, human
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Receptor-Interacting Protein Serine-Threonine Kinases
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MAP Kinase Kinase Kinases