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Chromophore packing leads to hysteresis in GFP.
Andrews BT, Roy M, Jennings PA. Andrews BT, et al. J Mol Biol. 2009 Sep 11;392(1):218-27. doi: 10.1016/j.jmb.2009.06.072. Epub 2009 Jul 3. J Mol Biol. 2009. PMID: 19577576 Free PMC article.