Melanargia galathea, the marbled white, is a medium-sized butterfly in the family Nymphalidae.[1] Despite its common name and appearance, this butterfly is one of the "browns", of the subfamily Satyrinae.

Marbled white
Male, upperside
Whitecross Green Wood, Oxfordshire, England
female, upperside
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Lepidoptera
Family: Nymphalidae
Genus: Melanargia
Species:
M. galathea
Binomial name
Melanargia galathea
Synonyms
  • Papilio galathea Linnaeus, 1758
  • Melanargia albibasa Lempke 1957
  • Melanargia annae Kesseler 1945
  • Melanargia brunneocosta Lempke 1957
  • Melanargia brunissime Perrier 1935
  • Melanargia caeca Lempke 1957
  • Melanargia carolina Weber 1964
  • Melanargia caronae Gallay 1947
  • Melanargia craskei Tubbs 1978
  • Melanargia dissiuncta Gussitsch 1917
  • Melanargia emma Rocci 1930
  • Agapetes essonsakaria Verity, 1953
  • Melanargia grisescens Varin 1948
  • Melanargia hades Reiss 1961
  • Melanargia laetepicta Stauder 1922
  • Melanargia lepigrei Betz 1948
  • Agapetes leucogonia Collier, 1952
  • Agapetes macronereus Verity, 1957
  • Agapetes macrosciritis Verity, 1953
  • Melanargia magnifica Stauder 1913
  • Agapetes microsciritis Verity, 1953
  • Satyrus nereine Verity, 1935
  • Melanargia nigrescens Varin 1948
  • Satyrus nigrionereus Verity, 1935
  • Melanargia pedemontii Rocci 1930
  • Agapetes pedenereus Verity, 1953
  • Agapetes pedenereine Verity, 1953
  • Melanargia planorum Rocci 1930
  • Melanargia pura Niepelt 1914
  • Melanargia quadriocellata Pionneau 1937
  • Melanargia sibillyna Rocci 1930
  • Melanargia sylvia Rocci 1930
  • Agapetes tenebrogigas Verity, 1938
  • Agapetes tenebronana Verity, 1938
  • Melanargia valentini Williams 1951
  • Melanargia vitrimontis Kesseler 1945

This species can be found across most of Europe, southern Russia, Asia Minor and Iran. It is found in forest clearings and edges, meadows and steppe where it occurs up to 1,500–1,700 metres (4,900–5,600 ft) above sea level. The caterpillars feed on various grasses.

Subspecies

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Subspecies include:[2]

  • Melanargia galathea galathea Europe, southern Urals
  • Melanargia galathea donsa Fruhstorfer, 1916 Caucasus
  • Melanargia galathea lucasi (Rambur, 1858) North Africa
  • Melanargia galathea magdalenae Reichl, 1975
  • Melanargia galathea procida (Herbst, 1796)
  • Melanargia galathea satnia Fruhstorfer, 1917 (= njurdzhan Sheljuzhko) Caucasus Major and Minor
  • Melanargia galathea tenebrosa Fruhstorfer, 1917


Distribution

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This species can be found across most of Europe, southern Russia, Asia Minor and Iran. There is an isolated population in Japan.[verification needed] It is not found in Ireland, North Britain, Scandinavia (except Denmark) and Portugal or Spain. The late twentieth century saw an expansion of its range in the UK.[3][4][2]

Habitat

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It is found in forest clearings and edges, meadows and steppe where it occurs up to 1,500-1,700 m above sea level.[4] They are a common sight in unimproved grasslands across southern Britain, particularly on the South Downs, but also extending slightly further north to places such as the Dunstable Downs, Devil's Dyke near Newmarket, Fleam Dyke, Les King Wood near the South Cambs village of Cottenham, The "Edwards" wood, Dry Drayton.

Description

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Illustration of egg, larva and pupa

Melanargia galathea has a wingspan of 46–56 millimetres (1.8–2.2 in).[5] In these medium-sized butterflies the upper side of the wings is decorated with white and gray-black or dark brown markings, but it is always gray-black or dark brown checkered in the basal and distal areas. The underside is similar to the upper side but the drawings is light gray or light brown. On the underside of the hindwings is present a row of gray eye spots. The males and the females are quite similar, except that some females may have a yellowish nuance on the underside of the wings. Seitz - M. galathea. In the otherwise black cell of both wing an oval white spot which is not divided by a transverse bar. On the hindwing above the ocelli are quite invisible or shine through very faintly from the underside.[6]


This species is rather similar to the Iberian marbled white (Melanargia lachesis) that replaces M. galathea in Spain and southern France.

The caterpillars are about 28 millimeters long. They are green or yellow with some lighter and darker narrow longitudinal lines. The head is always light brown.

Life cycle

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Like other members of its subfamily, the larvae feed on various grasses. These include Phleum (P. pratense), Poa (P. annua, P. trivalis), Festuca rubra, Bromus erectus, Dactylis, Brachypodium pinnatum, Agrostis capillaris, Elytrigia, Holcus, Dactylis, Triticum and Agropyron species .[2]

Eggs are laid on the wing, or from brief perches on grass stems, and are just sprinkled among the grass stems. Upon hatching, the larvae immediately enter hibernation and only feed the following spring when the fresh growth occurs. They are a lime-green colour, with a dark green line running down the middle of their back. Pupation takes place at ground level in a loose cocoon. Adults can be found from early June to early September.[4] On a good site, in warm, sunny weather, thousands can be seen gently fluttering amongst the grass heads.

Alternative common names

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The marbled white was called "Our Half-mourner" by James Petiver (1717), "The Marmoris" by Benjamin Wilkes and "The Marmoress" by Moses Harris.[citation needed]

References

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  1. ^ Catalogue of life
  2. ^ a b c "Melanargia Meigen, 1828" at Markku Savela's Lepidoptera and Some Other Life Forms
  3. ^ Fauna europaea
  4. ^ a b c Satyrinae of the Western Palearctic - Melanargia galathea
  5. ^ "European Butterfly Guide". Archived from the original on 2009-05-15. Retrieved 2017-08-07.
  6. ^ Seitz. A. in Seitz, A. ed. Band 1: Abt. 1, Die Großschmetterlinge des palaearktischen Faunengebietes, Die palaearktischen Tagfalter, 1909, 379 Seiten, mit 89 kolorierten Tafeln (3470 Figuren)  This article incorporates text from this source, which is in the public domain.
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